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downy rattlesnake-plantain, goodyérie pubescente

checkered rattlesnake-plantain, goodyérie panachée

Leaves

blade distinctly lined with white or greenish white on veins throughout, broadly elliptic to broadly ovate, 2.1–6.2 × 1.3–3 cm, apex acute or obtuse.

blade with lateral veins and sometimes midrib prominently to faintly bordered with bands of white, gray, or paler green tissue frequently merging, giving leaf rather blotched appearance, especially near margins and apex where veins are closer together, infrequently uniformly green or with only midrib whitened, narrowly elliptic to ovate, 1.4–5.5 × 0.9–2.6 cm, apex usually acute.

Inflorescences

10–57-flowered, cylindric (equally dense on all sides);

peduncle 11–35 cm.

densely to loosely spiraled or secund, 5–72-flowered;

peduncle 6–23 cm.

Flowers

lateral sepals 3.1–5.3 mm;

petals distinct;

hood 3.6–5.7 mm;

lip scrotiform, 2.5–4.2 × 2.2–3.5 mm, apex reflexed, outer surface slightly tuberculate, inner surface with somewhat thickened veins, without glandular processes;

anther inflexed, immersed in cup-shaped clinandrium, apex blunt;

pollinia blunt;

rostellum with marginal notch 0.1–0.4 mm deep;

viscidium orbiculate.

lateral sepals 3.8–6 mm;

petals connivent;

hood 3.9–7.1 mm;

lip deeply concave to saccate, 3–5.5 × 1.2–3.1 mm, apex spreading or recurved, blunt or acute, inner surface with 2 or 4 unequal rows of glandular papillae;

anther erect, base 0–1/3 immersed in shallowly concave to cup-shaped clinandrium, apex apiculate to acuminate;

pollinia acute or short-acuminate;

rostellar beak 2-pronged, 0.6–1.7 mm, equal to or longer than body of stigma;

viscidium elliptic.

2n

= 26.

= 45, 60.

Goodyera pubescens

Goodyera tesselata

Phenology Flowering mid Jul–early Sep. Flowering mid Jul–early Sep.
Habitat Almost any wooded habitat with acid surface conditions, most frequent on moist humus soils in shady, upland woods of hemlock, pine, oak, or maple, less frequent in lowland woods, bogs, swamps More common in dry or moist, upland, coniferous or mixed woods, less frequent in white-cedar swamps, margins of spruce-tamarack bogs
Elevation 0–1600 m (0–5200 ft) 0–300 m (0–1000 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WI; WV; NS; ON; QC
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from FNA
CT; MA; MD; ME; MI; MN; NH; NJ; NY; OH; PA; RI; VT; WI; MB; NB; NF; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Goodyera pubescens is very different in its lip and column morphology, phenolic constituents, and chromosome number (2n = 26) from other North American Goodyera species, indicating that it is not closely related to them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Goodyera tesselata is almost completely restricted to once-glaciated areas in the Great Lakes region, the northeastern United States, and adjacent Canada.

Goodyera tesselata tends to be taller and to have larger leaves, more cauline bracts, more flowers, and longer perianths than Goodyera repens. Although overlap exists in any count or measurement, the two are, for the most part, easily distinguishable. Some of the difficulty in identifying ambiguous specimens is explained by the hypothesis (J. A. Kallunki 1976) that G. tesselata itself is intermediate between and a probable allotetraploid of G. repens with white-reticulate leaves and G. oblongifolia. Because the tetraploids and the triploid hybrids (2n = ca. 45) cannot be distinguished with certainty by their morphology, the description here includes individuals of both tetraploid and triploid individuals. In the extreme, individuals of G. tesselata (in the broad sense) approach G. repens more often than G. oblongifolia, and it is sometimes impossible to distinguish some triploids from G. repens with white-reticulate leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 515. FNA vol. 26, p. 516.
Parent taxa Orchidaceae > subfam. Orchidoideae > tribe Cranichideae > subtribe Goodyerinae > Goodyera Orchidaceae > subfam. Orchidoideae > tribe Cranichideae > subtribe Goodyerinae > Goodyera
Sibling taxa
G. oblongifolia, G. repens, G. tesselata
G. oblongifolia, G. pubescens, G. repens
Synonyms Neottia pubescens, Peramium pubescens Peramium tesselatum
Name authority (Willdenow) R. Brown: in W. Aiton and W. T. Aiton, Hortus Kew. 5: 198. (1813) Loddiges: Bot. Cab. 10: plate 952. (1824)
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