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goodyérie, lattice-leaf, rattlesnake-plantain

Habit Herbs, terrestrial, rhizomatous, scapose, glabrous except for rather sticky, multicellular hairs on peduncles, bracts, sepals, and ovaries.
Roots

arising from nodes of rhizome, fibrous.

Stems

erect, with rosette of leaves, not succulent.

Leaves

evergreen, more than 1, in basal rosette, petiolate;

blade commonly marked with white to pale green.

Inflorescences

terminal, 5–72-flowered spikes, erect;

peduncles with sheathing bracts.

Flowers

resupinate, white, sometimes tinged green, ivory, or brown, sessile;

sepals distinct, nearly equal;

dorsal sepal and petals forming hood;

lip free from column, fleshy, base concave to saccate, apex ligulate or pointed;

anther 1, erect or inflexed;

pollinia 2, sectile;

rostellum notched or 2-pronged.

Fruits

capsules, erect, dehiscing along 3 ribs.

Goodyera

Distribution
from USDA
Nearly worldwide; primarily Southeast Asia; ca 16 species in Western Hemisphere
[BONAP county map]
Discussion

Species 40–100 (4 in the flora).

The four species of Goodyera in the flora are sometimes difficult to distinguish, especially without flowers. This difficulty is compounded, even with flowers, by the intermediate nature of Goodyera tesselata, which is likely an allotetraploid derived from G. repens with white-reticulate leaves and G. oblongifolia, and by the presence of triploid hybrids in some mixed populations of the three species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lip scrotiform, apex reflexed, no fleshy callosities on inner surface; rostellum notched; inflorescences cylindric (equally dense on all sides); leaves with midrib bordered by broad white bands and lateral veins traced with narrower white bands, both sharply demarcated from adjacent green tissue.
G. pubescens
1. Lip deeply concave or saccate, apex spreading or recurved, fleshy callosities on inner surface; rostellum with 2-pronged beak; inflorescences loosely spiraled or secund (infrequently cylindric); leaves uniformly green or reticulate with white or pale green on midrib and/or lateral veins.
→ 2
2. Leaf blades usually with only midrib whitened (infrequently with lateral veins, especially those near midrib, lightly penciled in white); sepals 5.7–7.8 mm; lip 4.9–7.9 mm; rostellar beak 2.3–3.6 mm; lip apex short, spreading or slightly arching with upright or involute margins.
G. oblongifolia
2. Leaf blades uniformly green or reticulate with white or pale green on lateral veins and sometimes midrib (very infrequently only midrib whitened in G. tesselata); sepals 3–6 mm; lip 1.8–5.5 mm; rostellar beak 1.7 mm or less; lip apex spreading or recurved with spreading margins.
→ 3
3. Lip narrowly saccate with elongate, recurved apex; rostellar beak 0.2–0.6 mm, shorter than body of stigma.
G. repens
3. Lip deeply concave with short spreading or recurved apex; rostellar beak 0.6–1.7 mm, equal to or longer than body of stigma.
G. tesselata
Source FNA vol. 26, p. 514. Author: Jacquelyn A. Kallunki.
Parent taxa Orchidaceae > subfam. Orchidoideae > tribe Cranichideae > subtribe Goodyerinae
Subordinate taxa
G. oblongifolia, G. pubescens, G. repens, G. tesselata
Name authority R. Brown: in W. Aiton and W. T. Aiton, Hortus Kew. 5: 197. (1813)
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