Asteraceae tribe Heliantheae subtribe Madiinae |
Lagophylla |
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hare-leaf, rabbitleaf |
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Habit | Annuals, perennials, subshrubs, or shrubs, (0.5–)1–250 cm. | Annuals, 8–100(–150) cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ± erect. |
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Leaves | mostly basal, or basal and cauline, or mostly cauline; opposite and/or alternate; usually sessile, rarely petiolate; blades usually spatulate to oblanceolate or linear to linear-elliptic, sometimes pinnately lobed, ultimate margins entire or toothed, faces usually hairy and glandular (glands sometimes elaborate, apices sometimes gland- or spine-tipped). |
mostly cauline; proximal opposite, most alternate; ± sessile; blades narrowly elliptic to linear or (proximal) oblanceolate to spatulate, margins usually entire (proximal sometimes toothed), faces hirsute to strigose, sericeous, or villous (all or distal sometimes stipitate-glandular as well). |
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Involucres | campanulate, cylindric, ellipsoid, globose, fusiform, hemispheric, obconic, or urceolate. |
± hemispheric or obovoid to obconic, 3–6+ mm diam. (sometimes subtended by calyculi of 2–5 bractlets). |
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Receptacles | flat or convex to conic, ± paleate (paleae either in 1 series interior to ray florets, then usually connate and persistent, or 1 palea subtending each disc floret, then usually distinct and soon falling, or, in discoid heads, paleae functioning as phyllaries in 1 series and together constituting an “involucre”). |
flat to convex, densely hirtellous, paleate (paleae in rings between rays and discs, distinct or proximally connate, scarious). |
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Ray florets | 0, or 1–75+, pistillate, fertile; corollas usually yellow or whitish, sometimes proximally yellow and distally whitish, often marked with red or purple or turning red on drying, rarely orange or red-orange (apices usually ± deeply divided, lobes usually 3, sometimes 2). |
5, pistillate, fertile; corollas yellow (often nerved with red to purple abaxially). |
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Disc florets | 1–200+, bisexual and fertile, or functionally staminate; corollas usually yellow, sometimes brown, orange, reddish, or whitish, tubes much shorter than to about equaling cylindric to funnelform throats, lobes 5, deltate; anther thecae usually dark (reddish to purple, often described as “black”), sometimes pale (yellow to brown); stigmatic papillae in 2 lines. |
6, functionally staminate; corollas yellow, tubes shorter than funnelform throats, lobes 5, deltate (anthers ± dark purple; styles glabrous proximal to branches). |
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Phyllaries | persistent or falling, 1–75+ in 1(–2) series or 0 (the “involucre” then interpreted as constituted of 1 series of receptacular paleae; phyllaries distinct or weakly connate, narrowly ovate to lanceolate or linear, equal or subequal, herbaceous, usually conduplicate, often each partly or wholly investing a subtended floret, apices often attenuate, faces usually glandular). |
5 in 1 series (linear to oblanceolate, herbaceous, each wholly enveloping a subtended ray ovary, abaxially piloso-hirsute to hirtellous or scabrellous). |
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Calyculi | usually 0, sometimes 2–12+ bractlets (Centromadia, some members of Hemizonia and Lagophylla). |
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Heads | usually radiate, rarely discoid, usually in corymbiform, glomerulate, paniculiform, spiciform, or ± umbelliform arrays, sometimes borne singly. |
radiate, in ± paniculiform arrays or in glomerules. |
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Cypselae | obcompressed, compressed, or terete, often obpyramidal, clavate, or fusiform, glabrous or hairy (ray and disc often different); pappi 0, or persistent or falling, of awns, bristles, or scales (sometimes in combination) in 1–2 series (the elements often ciliate or plumose). |
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Disc cypselae | 0; pappi 0. |
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Ray cypselae | ± obcompressed (attachments basal, apices beakless, faces glabrous); pappi 0. |
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Peduncular | bracts: pit-glands, tack-glands, and/or spines 0. |
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x | = 7. |
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Asteraceae tribe Heliantheae subtribe Madiinae |
Lagophylla |
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Distribution |
HI; South America; Mostly w North America |
w North America |
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Discussion | Genera 24, species 121 (20 genera, 83 species in the flora). Madiinae has long included the “tarweeds,” which are centered in or restricted to the California Floristic Province, the silversword alliance of the Hawaiian Islands (Argyroxiphium, Dubautia, and Wilkesia), and Raillardella (in a broad sense), which was formerly included in Senecioneae (Carlquist et al. 2003). Such a circumscription was used by H. Robinson (1981) and is followed here. In papers by B. G. Baldwin and B. Wessa (2000) and Baldwin et al. (2002), Madiinae were linked to genera traditionally included in Heliantheae (e.g., Eriophyllum, Hulsea, Lasthenia, Venegasia) and in Senecioneae (e.g., Arnica) and Madiinae plus Arnicinae, Baeriinae, Hulseinae, and Venegasiinae were included in tribe Madieae. Key to genera of Madiinae here is based on key by B. G. Baldwin (in S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 4 (4 in the flora). Molecular phylogenetic data have indicated that Lagophylla is sister to Layia (S. Carlquist et al. 2003). Members of Lagophylla are easily overlooked in the field; leaves usually wither before flowering and heads often close at midday. Ray laminae often shrivel greatly in pressed specimens (making lengths difficult to assess). The species are moderately interfertile (W. C. Thompson 1983); most do not co-occur and no natural hybrids have been documented. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 254. | FNA vol. 21, p. 260. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 390. (1841) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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