Asteraceae tribe Heliantheae subtribe Madiinae |
Centromadia |
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spikeweed, tarweed |
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Habit | Annuals, perennials, subshrubs, or shrubs, (0.5–)1–250 cm. | Annuals, 10–120 cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ± erect to prostrate. |
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Leaves | mostly basal, or basal and cauline, or mostly cauline; opposite and/or alternate; usually sessile, rarely petiolate; blades usually spatulate to oblanceolate or linear to linear-elliptic, sometimes pinnately lobed, ultimate margins entire or toothed, faces usually hairy and glandular (glands sometimes elaborate, apices sometimes gland- or spine-tipped). |
mostly cauline (at flowering); proximal opposite (often in winter–spring rosettes), most alternate; ± sessile; blades oblanceolate to linear or lance-linear, proximal usually 1–2-pinnatifid, ultimate margins toothed or entire (sometimes bristly-ciliate), (apices of distal leaves usually spine-tipped) faces glabrous, scabroso-hirtellous, ± hirsute, or villous, often glandular as well. |
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Involucres | campanulate, cylindric, ellipsoid, globose, fusiform, hemispheric, obconic, or urceolate. |
± obconic or urceolate, 3–8+ mm diam. (subtended by calyculi of 5–12+ usually spine-tipped bractlets). |
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Receptacles | flat or convex to conic, ± paleate (paleae either in 1 series interior to ray florets, then usually connate and persistent, or 1 palea subtending each disc floret, then usually distinct and soon falling, or, in discoid heads, paleae functioning as phyllaries in 1 series and together constituting an “involucre”). |
flat to convex, setulose, paleate (paleae persistent, subtending all or most disc florets, distinct, phyllary-like, more scarious). |
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Ray florets | 0, or 1–75+, pistillate, fertile; corollas usually yellow or whitish, sometimes proximally yellow and distally whitish, often marked with red or purple or turning red on drying, rarely orange or red-orange (apices usually ± deeply divided, lobes usually 3, sometimes 2). |
5–75+, pistillate, fertile; corollas yellow. |
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Disc florets | 1–200+, bisexual and fertile, or functionally staminate; corollas usually yellow, sometimes brown, orange, reddish, or whitish, tubes much shorter than to about equaling cylindric to funnelform throats, lobes 5, deltate; anther thecae usually dark (reddish to purple, often described as “black”), sometimes pale (yellow to brown); stigmatic papillae in 2 lines. |
6–200+, usually functionally staminate, rarely bisexual and fertile; corollas yellow, tubes shorter than funnelform throats, lobes 5, deltate (anthers reddish to dark purple or yellow to brownish; styles glabrous proximal to branches). |
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Phyllaries | persistent or falling, 1–75+ in 1(–2) series or 0 (the “involucre” then interpreted as constituted of 1 series of receptacular paleae; phyllaries distinct or weakly connate, narrowly ovate to lanceolate or linear, equal or subequal, herbaceous, usually conduplicate, often each partly or wholly investing a subtended floret, apices often attenuate, faces usually glandular). |
falling or persistent, 5–75+ in 1 series (lanceolate to lance-attenuate or oblanceolate, herbaceous, each usually 1/2 enveloping subtended ray floret proximally, abaxially scabroso-hirtellous, hirsute, or villous and/or glandular, apices often spine-tipped). |
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Calyculi | usually 0, sometimes 2–12+ bractlets (Centromadia, some members of Hemizonia and Lagophylla). |
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Heads | usually radiate, rarely discoid, usually in corymbiform, glomerulate, paniculiform, spiciform, or ± umbelliform arrays, sometimes borne singly. |
radiate, borne in glomerules or ± spiciform-paniculiform or ± umbelliform arrays. |
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Cypselae | obcompressed, compressed, or terete, often obpyramidal, clavate, or fusiform, glabrous or hairy (ray and disc often different); pappi 0, or persistent or falling, of awns, bristles, or scales (sometimes in combination) in 1–2 series (the elements often ciliate or plumose). |
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Disc cypselae | usually 0; pappi (of disc florets) 0 or of 3–12 linear, oblanceolate, or subulate scales. |
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Ray cypselae | ± compressed (abaxially gibbous, basal attachments basal or oblique, apices beaked or elevated adaxially, faces glabrous); pappi 0. |
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Peduncular | bracts: pit-glands and tack-glands 0 (apices usually spine-tipped, sometimes apiculate). |
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x | = 13. |
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Asteraceae tribe Heliantheae subtribe Madiinae |
Centromadia |
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Distribution |
HI; South America; Mostly w North America |
w United States; nw Mexico |
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Discussion | Genera 24, species 121 (20 genera, 83 species in the flora). Madiinae has long included the “tarweeds,” which are centered in or restricted to the California Floristic Province, the silversword alliance of the Hawaiian Islands (Argyroxiphium, Dubautia, and Wilkesia), and Raillardella (in a broad sense), which was formerly included in Senecioneae (Carlquist et al. 2003). Such a circumscription was used by H. Robinson (1981) and is followed here. In papers by B. G. Baldwin and B. Wessa (2000) and Baldwin et al. (2002), Madiinae were linked to genera traditionally included in Heliantheae (e.g., Eriophyllum, Hulsea, Lasthenia, Venegasia) and in Senecioneae (e.g., Arnica) and Madiinae plus Arnicinae, Baeriinae, Hulseinae, and Venegasiinae were included in tribe Madieae. Key to genera of Madiinae here is based on key by B. G. Baldwin (in S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 4 (3 in the flora). Following B. G. Baldwin (1999b), Centromadia is treated here as distinct from Hemizonia; Centromadia is more closely related to Calycadenia, Deinandra, Holocarpha, and Osmadenia than to Hemizonia in the strict sense (S. Carlquist et al. 2003). Taxa of Centromadia are self-incompatible and of low to high interfertility (C. S. Venkatesh 1958). Most occur in somewhat poorly drained or alkaline sites. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 254. | FNA vol. 21, p. 276. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) | Greene: Fl. Francisc. 4: 424. (1897) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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