Asteraceae tribe Heliantheae subtribe Madiinae |
Blepharipappus |
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blepharipappus, eyelash tarweed, rough eyelashweed |
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Habit | Annuals, perennials, subshrubs, or shrubs, (0.5–)1–250 cm. | Annuals, 5–20(–40+) cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | ± erect (scabrous and, sometimes, hirsute, usually stipitate-glandular distally). |
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Leaves | mostly basal, or basal and cauline, or mostly cauline; opposite and/or alternate; usually sessile, rarely petiolate; blades usually spatulate to oblanceolate or linear to linear-elliptic, sometimes pinnately lobed, ultimate margins entire or toothed, faces usually hairy and glandular (glands sometimes elaborate, apices sometimes gland- or spine-tipped). |
mostly cauline; proximal opposite, most alternate; sessile; blades narrowly spatulate to linear, margins entire, faces scabrous, hirsute, strigose, sericeous, or villous (distal leaves usually stipitate-glandular as well). |
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Involucres | campanulate, cylindric, ellipsoid, globose, fusiform, hemispheric, obconic, or urceolate. |
turbinate to campanulate or hemispheric, 3–6+ mm diam. |
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Receptacles | flat or convex to conic, ± paleate (paleae either in 1 series interior to ray florets, then usually connate and persistent, or 1 palea subtending each disc floret, then usually distinct and soon falling, or, in discoid heads, paleae functioning as phyllaries in 1 series and together constituting an “involucre”). |
convex, glabrous, paleate (paleae falling, subtending all or most disc florets, outer herbaceous, inner scarious). |
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Ray florets | 0, or 1–75+, pistillate, fertile; corollas usually yellow or whitish, sometimes proximally yellow and distally whitish, often marked with red or purple or turning red on drying, rarely orange or red-orange (apices usually ± deeply divided, lobes usually 3, sometimes 2). |
(2–)3–5(–8), pistillate, fertile; corollas whitish (nerved with purple abaxially). |
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Disc florets | 1–200+, bisexual and fertile, or functionally staminate; corollas usually yellow, sometimes brown, orange, reddish, or whitish, tubes much shorter than to about equaling cylindric to funnelform throats, lobes 5, deltate; anther thecae usually dark (reddish to purple, often described as “black”), sometimes pale (yellow to brown); stigmatic papillae in 2 lines. |
6–25(–60+), bisexual, fertile; corollas whitish, tubes shorter than or about equaling funnelform throats, lobes 5, deltate (anthers ± dark purple; styles ± hairy proximal to branches, branches ca. 0.2 mm). |
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Phyllaries | persistent or falling, 1–75+ in 1(–2) series or 0 (the “involucre” then interpreted as constituted of 1 series of receptacular paleae; phyllaries distinct or weakly connate, narrowly ovate to lanceolate or linear, equal or subequal, herbaceous, usually conduplicate, often each partly or wholly investing a subtended floret, apices often attenuate, faces usually glandular). |
(2–)3–5(–8) in 1 series, ± lanceolate or oblanceolate, herbaceous, each ± 1/2 investing subtended floret proximally, abaxially ± hirsute and/or stipitate-glandular. |
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Calyculi | usually 0, sometimes 2–12+ bractlets (Centromadia, some members of Hemizonia and Lagophylla). |
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Heads | usually radiate, rarely discoid, usually in corymbiform, glomerulate, paniculiform, spiciform, or ± umbelliform arrays, sometimes borne singly. |
radiate, borne singly or in open, corymbiform arrays. |
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Cypselae | obcompressed, compressed, or terete, often obpyramidal, clavate, or fusiform, glabrous or hairy (ray and disc often different); pappi 0, or persistent or falling, of awns, bristles, or scales (sometimes in combination) in 1–2 series (the elements often ciliate or plumose). |
± obconic or terete (basal attachments ± central, faces ± villous, apices not beaked); pappi 0 or of 12–18(–26) subulate fimbriate to ciliate or plumose scales. |
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Peduncular | bracts: pit-glands, tack-glands, and/or spines 0 at tips. |
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x | = 8. |
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Asteraceae tribe Heliantheae subtribe Madiinae |
Blepharipappus |
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Distribution |
HI; South America; Mostly w North America |
nw North America |
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Discussion | Genera 24, species 121 (20 genera, 83 species in the flora). Madiinae has long included the “tarweeds,” which are centered in or restricted to the California Floristic Province, the silversword alliance of the Hawaiian Islands (Argyroxiphium, Dubautia, and Wilkesia), and Raillardella (in a broad sense), which was formerly included in Senecioneae (Carlquist et al. 2003). Such a circumscription was used by H. Robinson (1981) and is followed here. In papers by B. G. Baldwin and B. Wessa (2000) and Baldwin et al. (2002), Madiinae were linked to genera traditionally included in Heliantheae (e.g., Eriophyllum, Hulsea, Lasthenia, Venegasia) and in Senecioneae (e.g., Arnica) and Madiinae plus Arnicinae, Baeriinae, Hulseinae, and Venegasiinae were included in tribe Madieae. Key to genera of Madiinae here is based on key by B. G. Baldwin (in S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 1. Blepharipappus and members of Layia were once treated as congeneric; Layia appears to be more closely related to Lagophylla than to Blepharipappus, based on molecular phylogenetic data (S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 254. | FNA vol. 21, p. 259. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) | Hooker: Fl. Bor.-Amer. 1: 316. (1833) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |