Asteraceae tribe Heliantheae subtribe Madiinae |
Achyrachaena |
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blow-wives |
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Habit | Annuals, perennials, subshrubs, or shrubs, (0.5–)1–250 cm. | Annuals, 4–62 cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, branched distally. |
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Leaves | mostly basal, or basal and cauline, or mostly cauline; opposite and/or alternate; usually sessile, rarely petiolate; blades usually spatulate to oblanceolate or linear to linear-elliptic, sometimes pinnately lobed, ultimate margins entire or toothed, faces usually hairy and glandular (glands sometimes elaborate, apices sometimes gland- or spine-tipped). |
mostly cauline; proximal opposite, distal alternate; ± sessile; blades linear, margins entire or toothed, faces hirsute or villous and (distal leaves) sparsely glandular-pubescent. |
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Involucres | campanulate, cylindric, ellipsoid, globose, fusiform, hemispheric, obconic, or urceolate. |
± campanulate to cylindric or ellipsoid, 3–12+ mm diam. |
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Receptacles | flat or convex to conic, ± paleate (paleae either in 1 series interior to ray florets, then usually connate and persistent, or 1 palea subtending each disc floret, then usually distinct and soon falling, or, in discoid heads, paleae functioning as phyllaries in 1 series and together constituting an “involucre”). |
flat to convex (setulose), paleate (paleae falling, in 1 series between rays and disc, distinct). |
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Ray florets | 0, or 1–75+, pistillate, fertile; corollas usually yellow or whitish, sometimes proximally yellow and distally whitish, often marked with red or purple or turning red on drying, rarely orange or red-orange (apices usually ± deeply divided, lobes usually 3, sometimes 2). |
3–8, pistillate, fertile; corollas yellow, turning reddish (lobes ± parallel). |
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Disc florets | 1–200+, bisexual and fertile, or functionally staminate; corollas usually yellow, sometimes brown, orange, reddish, or whitish, tubes much shorter than to about equaling cylindric to funnelform throats, lobes 5, deltate; anther thecae usually dark (reddish to purple, often described as “black”), sometimes pale (yellow to brown); stigmatic papillae in 2 lines. |
4–35, bisexual and fertile; corollas yellow to reddish, tubes ± equaling narrowly funnelform throats, lobes 5, lance-deltate (anthers ± dark purple; styles glabrous proximal to branches). |
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Phyllaries | persistent or falling, 1–75+ in 1(–2) series or 0 (the “involucre” then interpreted as constituted of 1 series of receptacular paleae; phyllaries distinct or weakly connate, narrowly ovate to lanceolate or linear, equal or subequal, herbaceous, usually conduplicate, often each partly or wholly investing a subtended floret, apices often attenuate, faces usually glandular). |
3–8 in 1 series (± lance-linear, herbaceous, each fully enveloping a ray ovary, abaxially hirsute or villous and sparsely glandular-pubescent or eglandular). |
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Calyculi | usually 0, sometimes 2–12+ bractlets (Centromadia, some members of Hemizonia and Lagophylla). |
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Heads | usually radiate, rarely discoid, usually in corymbiform, glomerulate, paniculiform, spiciform, or ± umbelliform arrays, sometimes borne singly. |
radiate, borne singly or in loose, ± corymbiform arrays. |
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Cypselae | obcompressed, compressed, or terete, often obpyramidal, clavate, or fusiform, glabrous or hairy (ray and disc often different); pappi 0, or persistent or falling, of awns, bristles, or scales (sometimes in combination) in 1–2 series (the elements often ciliate or plumose). |
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Disc cypselae | (brown to black) similar to rays (± scabrous); pappi of 10 white, oblong scales (apices obtuse). |
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Ray cypselae | (black) obcompressed, clavate (10-ribbed, straight, basal attachments centered, apices beakless, faces glabrous or ± scabrous); pappi 0. |
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Peduncular | bracts: pit-glands, tack-glands, and/or spines 0 at tips. |
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x | = 8. |
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Asteraceae tribe Heliantheae subtribe Madiinae |
Achyrachaena |
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Distribution |
HI; South America; Mostly w North America |
w North America; nw Mexico |
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Discussion | Genera 24, species 121 (20 genera, 83 species in the flora). Madiinae has long included the “tarweeds,” which are centered in or restricted to the California Floristic Province, the silversword alliance of the Hawaiian Islands (Argyroxiphium, Dubautia, and Wilkesia), and Raillardella (in a broad sense), which was formerly included in Senecioneae (Carlquist et al. 2003). Such a circumscription was used by H. Robinson (1981) and is followed here. In papers by B. G. Baldwin and B. Wessa (2000) and Baldwin et al. (2002), Madiinae were linked to genera traditionally included in Heliantheae (e.g., Eriophyllum, Hulsea, Lasthenia, Venegasia) and in Senecioneae (e.g., Arnica) and Madiinae plus Arnicinae, Baeriinae, Hulseinae, and Venegasiinae were included in tribe Madieae. Key to genera of Madiinae here is based on key by B. G. Baldwin (in S. Carlquist et al. 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 1. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 254. | FNA vol. 21, p. 258. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) | Schauer: Index Seminum (Bratislave) 1837: [3]. (1837) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |