Xanthium |
Xanthium strumarium |
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cocklebur, lampourde |
Canada cocklebur, cocklebur, common cocklebur, lampourde glouteron, rough cocklebur, rough cockleburr |
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Habit | Annuals (coarse), 10–200+ cm. | Plants 10–80(–200) cm; nodal spines 0. | ||||
Stems | erect, branched. |
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Leaves | cauline; mostly alternate (proximal 2–6 sometimes opposite); petiolate; blades lanceolate, linear, ovate, rounded-deltate, or suborbiculate, often ± palmately or pinnately lobed, ultimate margins entire or ± toothed, faces hirtellous or ± strigose, usually gland-dotted as well. |
petioles 20–100(–140+) mm; blades suborbiculate to ± pentagonal or deltate, 4–12(–18+) × 3–10(–18+) cm, sometimes palmately 3–5-lobed, abaxial faces green, hirtellous. |
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Staminate heads | involucres ± saucer-shaped, 3–5 mm diam.; phyllaries 6–16+ in 1–2+ series, distinct to bases; receptacles conic to columnar; paleae spatulate to cuneiform or linear, membranous, distally ± villous or hirtellous; florets 20–50+, corollas whitish, ± funnelform, lobes 5, erect or reflexed (filaments connate, anthers distinct or weakly coherent). |
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Pistillate heads | involucres ± ellipsoid, 2–5+ mm diam. at anthesis (6–20+ mm diam. at maturity); phyllaries 30–75+ in 6–12+ series, outer 5–8 distinct, the rest (sometimes interpreted as paleae) proximally connate, their distinct tips mostly ± hooked (the distal 1–3 usually longer, stouter, and not hooked), the whole becoming a hard, prickly perigynium (a bur); florets 2, corollas 0. |
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Heads | discoid, either pistillate (proximal) or functionally staminate (distal), in racemiform to spiciform arrays or borne singly (in axils). |
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Cypselae | (black) ± fusiform, enclosed in obovoid to ellipsoid, hard, prickly, 2-chambered burs; pappi 0. |
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Burs | 10–30+ mm. |
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x | = 18. |
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2n | = 36. |
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Xanthium |
Xanthium strumarium |
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Phenology | Flowering Jul–Oct. | |||||
Habitat | Damp or seasonally wet, often alkaline, soils, waste places, margins of agriculture | |||||
Elevation | 10–2000 m (0–6600 ft) | |||||
Distribution |
New World; introduced nearly worldwide |
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico; Central America; South America [Widely introduced in Old World]
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Discussion | Species 2–3 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Recognition of a dozen or more taxa (treated as species, subspecies, varieties, and/or forms) has been proposed for plants treated together here as Xanthium strumarium. Bases for the various taxa mostly involved subtle differences in the burs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 19. | FNA vol. 21, p. 19. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | X. americanum, X. chasei, X. chinense, X. curvescens, X. cylindricum, X. echinatum, X. echinellum, X. globosum, X. inflexum, X. italicum, X. orientale, X. oviforme, X. pensylvanicum, X. speciosum, X. strumarium var. canadense, X. strumarium var. glabratum, X. varians, X. wootonii | |||||
Name authority | Linnaeus: Sp. Pl. 2: 987. (1753): Gen. Pl. ed. 5, 424. (1754) | Linnaeus: Sp. Pl. 2: 987. (1753) | ||||
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