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cocklebur, lampourde

Canada cocklebur, cocklebur, common cocklebur, lampourde glouteron, rough cocklebur, rough cockleburr

Habit Annuals (coarse), 10–200+ cm. Plants 10–80(–200) cm; nodal spines 0.
Stems

erect, branched.

Leaves

cauline; mostly alternate (proximal 2–6 sometimes opposite);

petiolate;

blades lanceolate, linear, ovate, rounded-deltate, or suborbiculate, often ± palmately or pinnately lobed, ultimate margins entire or ± toothed, faces hirtellous or ± strigose, usually gland-dotted as well.

petioles 20–100(–140+) mm;

blades suborbiculate to ± pentagonal or deltate, 4–12(–18+) × 3–10(–18+) cm, sometimes palmately 3–5-lobed, abaxial faces green, hirtellous.

Staminate heads

involucres ± saucer-shaped, 3–5 mm diam.;

phyllaries 6–16+ in 1–2+ series, distinct to bases;

receptacles conic to columnar;

paleae spatulate to cuneiform or linear, membranous, distally ± villous or hirtellous;

florets 20–50+, corollas whitish, ± funnelform, lobes 5, erect or reflexed (filaments connate, anthers distinct or weakly coherent).

Pistillate heads

involucres ± ellipsoid, 2–5+ mm diam. at anthesis (6–20+ mm diam. at maturity);

phyllaries 30–75+ in 6–12+ series, outer 5–8 distinct, the rest (sometimes interpreted as paleae) proximally connate, their distinct tips mostly ± hooked (the distal 1–3 usually longer, stouter, and not hooked), the whole becoming a hard, prickly perigynium (a bur);

florets 2, corollas 0.

Heads

discoid, either pistillate (proximal) or functionally staminate (distal), in racemiform to spiciform arrays or borne singly (in axils).

Cypselae

(black) ± fusiform, enclosed in obovoid to ellipsoid, hard, prickly, 2-chambered burs;

pappi 0.

Burs

10–30+ mm.

x

= 18.

2n

= 36.

Xanthium

Xanthium strumarium

Phenology Flowering Jul–Oct.
Habitat Damp or seasonally wet, often alkaline, soils, waste places, margins of agriculture
Elevation 10–2000 m (0–6600 ft)
Distribution
from USDA
New World; introduced nearly worldwide
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico; Central America; South America [Widely introduced in Old World]
[WildflowerSearch map]
[BONAP county map]
Discussion

Species 2–3 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Recognition of a dozen or more taxa (treated as species, subspecies, varieties, and/or forms) has been proposed for plants treated together here as Xanthium strumarium. Bases for the various taxa mostly involved subtle differences in the burs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Nodal spines 0; leaf blades suborbiculate to ± pentagonal or deltate
X. strumarium
1. Nodal spines (1–)3-lobed, 15–30+ mm; leaf blades ± lanceolate to ovate or lance-linear
X. spinosum
Source FNA vol. 21, p. 19. Author: John L. Strother. FNA vol. 21, p. 19.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Ambrosiinae Asteraceae > tribe Heliantheae > subtribe Ambrosiinae > Xanthium
Sibling taxa
X. spinosum
Subordinate taxa
X. spinosum, X. strumarium
Synonyms X. americanum, X. chasei, X. chinense, X. curvescens, X. cylindricum, X. echinatum, X. echinellum, X. globosum, X. inflexum, X. italicum, X. orientale, X. oviforme, X. pensylvanicum, X. speciosum, X. strumarium var. canadense, X. strumarium var. glabratum, X. varians, X. wootonii
Name authority Linnaeus: Sp. Pl. 2: 987. (1753): Gen. Pl. ed. 5, 424. (1754) Linnaeus: Sp. Pl. 2: 987. (1753)
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