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barren-strawberry, waldsteinie

Habit Herbs, perennial, scapose, often mat-forming, 1–4 dm, strigose; rhizomatous. Herbs, shrubs, or subshrubs.
Stems

1–20+, ± erect.

Leaves

deciduous, basal, sometimes clustered around scape, simple or compound;

stipules adnate to petiole base, narrow, scarious, margins entire;

petiole present;

blade ± rounded, deeply cordate, ternate or 3–5(–7)-lobed, 3–5 cm, herbaceous, margins flat, when lobes usually shallow, dentate, often with large teeth distally, venation palmate, surfaces strigose or hirsute.

alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound;

stipules present, rarely absent.

Inflorescences

terminal, 3–8-flowered, panicles, open, ± as long as petioles, elongating somewhat in fruit;

bracts present;

bracteoles absent.

Pedicels

present.

Flowers

8–20 mm diam.;

epicalyx bractlets 0 [5, minute];

hypanthium broadly obconic to obcampanulate, slightly contracted at mouth, 2–4 mm, strigose or glabrous;

sepals 5, spreading, triangular;

petals 5, yellow, obovate to elliptic, elliptic-lanceolate, or suborbiculate;

stamens 50–70, ± equal to petals, filaments persistent;

torus not evident, receptacle base villous;

carpels 3–7, short-villous;

ovule 1.

torus usually enlarged, sometimes small or absent;

carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae);

ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula).

Fruits

aggregated achenes, 3–7, broadly crescent-shaped, 1.5–2.5 mm, short-villous;

hypanthium persistent;

sepals persistent, ± reflexed;

styles deciduous, not hooked.

achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets;

styles persistent or deciduous, not elongate (elongate but not plumose in Geum).

x

= 7.

= 7(8).

Waldsteinia

Rosaceae subfam. rosoideae

Distribution
from USDA
North America; Eurasia; temperate regions
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
Discussion

Species 6 or 7 (4 in the flora).

Plants of Waldsteinia are distinctive, small, rhizomatous, and often mat-forming herbs of woodlands with ternate or three- to five-lobed leaves, differing from Geum by their long, straight, deciduous styles. J. E. E. Smedmark (2006) showed that the type species of Waldsteinia is nested in Geum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily.

Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves mostly ternately compound, rarely merely deeply lobed
→ 2
1. Leaves 3–5(–7)-lobed, shallowly so or to ± 2/3 cleft
→ 3
2. Petals obovate to broadly elliptic, 3.5–10 mm, much longer than sepals.
W. fragarioides
2. Petals elliptic to elliptic-lanceolate, 2.5–4.5 mm, shorter than to barely exceeding sepals.
W. parviflora
3. Flowers 8–10 mm diam.; petals narrowly elliptic, barely exceeding sepals, bright yellow.
W. lobata
3. Flowers 15–18 mm diam.; petals broadly elliptic to suborbiculate, much longer than sepals, creamy yellow.
W. idahoensis
Source FNA vol. 9, p. 71. Author: James B. Phipps. FNA vol. 9, p. 23. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Rosoideae > tribe Colurieae Rosaceae
Subordinate taxa
W. fragarioides, W. idahoensis, W. lobata, W. parviflora
Name authority Willdenow: Ges. Naturf. Freunde Berlin Neue Schriften 2: 105, plate 4, fig. 1. (1799) Arnott: Botany, 107. (1832)
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