Waldsteinia |
Rosaceae subfam. rosoideae |
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barren-strawberry, waldsteinie |
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Habit | Herbs, perennial, scapose, often mat-forming, 1–4 dm, strigose; rhizomatous. | Herbs, shrubs, or subshrubs. | ||||||||||||
Stems | 1–20+, ± erect. |
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Leaves | deciduous, basal, sometimes clustered around scape, simple or compound; stipules adnate to petiole base, narrow, scarious, margins entire; petiole present; blade ± rounded, deeply cordate, ternate or 3–5(–7)-lobed, 3–5 cm, herbaceous, margins flat, when lobes usually shallow, dentate, often with large teeth distally, venation palmate, surfaces strigose or hirsute. |
alternate, rarely opposite, pinnately compound, sometimes simple or palmately compound; stipules present, rarely absent. |
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Inflorescences | terminal, 3–8-flowered, panicles, open, ± as long as petioles, elongating somewhat in fruit; bracts present; bracteoles absent. |
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Pedicels | present. |
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Flowers | 8–20 mm diam.; epicalyx bractlets 0 [5, minute]; hypanthium broadly obconic to obcampanulate, slightly contracted at mouth, 2–4 mm, strigose or glabrous; sepals 5, spreading, triangular; petals 5, yellow, obovate to elliptic, elliptic-lanceolate, or suborbiculate; stamens 50–70, ± equal to petals, filaments persistent; torus not evident, receptacle base villous; carpels 3–7, short-villous; ovule 1. |
torus usually enlarged, sometimes small or absent; carpels 1–260(–450), distinct, free, styles distinct, rarely connate (Roseae); ovules 1(or 2), collateral (Rubeae) or superposed (Fallugia, Filipendula). |
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Fruits | aggregated achenes, 3–7, broadly crescent-shaped, 1.5–2.5 mm, short-villous; hypanthium persistent; sepals persistent, ± reflexed; styles deciduous, not hooked. |
achenes or aggregated achenes sometimes with fleshy, urn-shaped hypanthium or enlarged torus, sometimes aggregated drupelets; styles persistent or deciduous, not elongate (elongate but not plumose in Geum). |
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x | = 7. |
= 7(8). |
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Waldsteinia |
Rosaceae subfam. rosoideae |
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Distribution |
North America; Eurasia; temperate regions |
North America; Mexico; Central America; South America; West Indies; Bermuda; Eurasia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia |
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Discussion | Species 6 or 7 (4 in the flora). Plants of Waldsteinia are distinctive, small, rhizomatous, and often mat-forming herbs of woodlands with ternate or three- to five-lobed leaves, differing from Geum by their long, straight, deciduous styles. J. E. E. Smedmark (2006) showed that the type species of Waldsteinia is nested in Geum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Variation in the number of genera in subfam. Rosoideae is due to differences in generic delimitation between D. Potter et al. (2007) and the authors of some Potentilleae genera. Cyanogenic glycosides and sorbitol are absent in the subfamily. Tribes 6, genera 28–35, species ca. 1600 (6 tribes, 26 genera, 302 species, including 1 hybrid, in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 71. | FNA vol. 9, p. 23. | ||||||||||||
Parent taxa | ||||||||||||||
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Name authority | Willdenow: Ges. Naturf. Freunde Berlin Neue Schriften 2: 105, plate 4, fig. 1. (1799) | Arnott: Botany, 107. (1832) | ||||||||||||
Web links |