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tonella

Habit Herbs, annual.
Stems

ascending to erect, glabrous proximally, glandular-hairy distally.

Leaves

cauline, opposite;

petiole present or absent;

blade not fleshy, not leathery, margins entire, crenate, dentate, or serrate.

Inflorescences

terminal, racemes;

bracts present.

Pedicels

present, glandular-hairy, sometimes glabrous or glabrate;

bracteoles absent.

Flowers

bisexual;

sepals 5, proximally connate, calyx ± bilaterally symmetric, subrotate to campanulate, lobes triangular to lanceolate;

corolla white on tube and lobes proximally, blue to violet or lavender on lobes distally, with dark violet spots internally near bases of adaxial and lateral lobes, bilaterally symmetric, weakly bilabiate, subrotate, tube base gibbous adaxially, sometimes obscurely so, not spurred abaxially, throat not densely pilose internally, lobes 5, abaxial 3, middle lobe of abaxial lip not folded lengthwise, not enclosing stamens and style, adaxial 2;

stamens 4, medially adnate to corolla, equal or ± didynamous, exserted, filaments glandular;

staminode 1, minute;

ovary 2-locular, placentation axile;

stigma linear.

Fruits

capsules, dehiscence septicidal and loculicidal.

Seeds

2 or 4, dark brown to black, reniform to ovoid, wings absent.

Tonella

Distribution
from USDA
w North America
[BONAP county map]
Discussion

Species 2 (2 in the flora).

Studies of vascular anatomy, organography, and gametophyte development (G. F. Schrock and B. F. Palser 1967) and molecular phylogenetic analyses (B. G. Baldwin et al. 2011) support the monophyly of Tonella and its sister-relationship to Collinsia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Corollas 5–7 × 6–12 mm; racemes: flowers 2–10 per node.
T. floribunda
1. Corollas 2–2.5 × 2–4 mm; racemes: flowers 1–3 per node.
T. tenella
Source FNA vol. 17, p. 256. Author: Craig C. Freeman.
Parent taxa Plantaginaceae
Subordinate taxa
T. floribunda, T. tenella
Name authority Nuttall ex A. Gray: Proc. Amer. Acad. Arts 7: 378. (1868)
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