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globe-mallow

hot springs globemallow

Habit Herbs, annual or perennial. Plants perennial.
Stems

erect or decumbent to ascending, stellate-canescent to stellate-silvery-lepidote, rarely glabrous.

erect, white or yellow, to 10–20 dm, rubbery, densely soft stellate-pubescent.

Leaves

petiolate or sessile;

stipules persistent or deciduous, linear;

blade linear, lanceolate, orbiculate, or ovate to triangular, trullate, or cordate, unlobed or pedately dissected, base tapered, cuneate, or truncate to cordate, margins entire or crenate to serrate.

blades white or yellow, deltate to lanceolate, subhastate to 3-lobed, 4–7 cm, secondary lobes to 2.2 cm, not rugose, base cuneate, margins crenate to dentate, surfaces green- to white-canescent.

Inflorescences

terminal, racemes or panicles, or flowers sometimes fascicled or in axillary cymose racemes;

involucel absent or present, involucellar bractlets inflated, cylindric, ovoid, flattened-spheric, flattened-spheric-conic, hemispheric, widely conic, truncate-conic, spheric, truncate-ovoid, ellipsoid, short-urceolate, helmet-shaped, or truncate-spheric, indurate, hairy;

mericarps [7–]9–30, drying tan to brown, 1- or 2-celled, without dorsal spur, distally extended or not, usually smooth, apex cuspidate, truncate, or acute, with or without seeds, proximally usually rugose-reticulate and indehiscent, distally ± dehiscent, often remaining attached to fruit axis after maturity by threadlike extension of dorsal vein.

paniculate, open, many-flowered, interrupted, tip leafy;

involucellar bractlets green to tan.

Flowers

sepals 6–7 mm;

petals white, pink, lavender, purple, red-orange, or red, 10–13 mm;

anthers yellow.

Seeds

1 or 2(or 3) per mericarp, glabrous or slightly pubescent.

1 or 2 per mericarp, gray, pubescent.

Schizocarps

short-urceolate;

mericarps 12–14, (3.5–)4–5.5 × 2–3 mm, chartaceous, nonreticulate dehiscent part 60% of height, with usually reflexed cusps to 2 mm, indehiscent part not wider than dehiscent part.

x

= 5.

2n

=20.

Sphaeralcea

Sphaeralcea polychroma

Phenology Flowering summer.
Habitat Desert lowlands
Elevation 1900 m (6200 ft)
Distribution
from USDA
w North America; Mexico; c North America; South America (Argentina, Bolivia, Brazil, Chile, Paraguay)
[BONAP county map]
from FNA
AZ; NM; TX; Mexico (Chihuahua)
[BONAP county map]
Discussion

Species ca. 50 (26 in the flora).

Sphaeralcea is often collected and somewhat difficult to identify. The species boundaries are not always sharp because there is frequent hybridization, polyploidy, and morphological variation in response to environmental conditions, particularly rainfall. Plants growing in Arizona are particularly difficult to identify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sphaeralcea polychroma is frequent in central New Mexico and western Texas. It is closely related to S. procera.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants annual, biennial, or perennial; mericarps: dehiscent part 10–75% of height, indehiscent part ± equaling to usually wider than dehiscent part
→ 2
1. Plants perennial; mericarps: dehiscent part 50–85% of height, indehiscent part usually not notably wider than dehiscent part
→ 9
2. Leaf blade surfaces silvery stellate-lepidote.
S. leptophylla
2. Leaf blade surfaces stellate-pubescent or stellate-canescent
→ 3
3. Plants annual; mericarps 1.5–2 mm; leaf blades gray-green pubescent, unlobed or 3–5-lobed.
S. coulteri
3. Plants usually perennial, sometimes annual or biennial; mericarps 2–5.5 mm; leaf blades green, pale green, yellow, yellow-green, or gray, light gray, or gray-green pubescent, unlobed, or if green or gray to green pubescent, leaves unlobed, 3–5-lobed, or pedately divided
→ 4
4. Leaf blades trullate or triangular, not deeply divided.
S. orcuttii
5. Stems 1–3(–5) dm
→ 7
6. Sepals 5–13 mm; petals 15–20(–25) mm
S. gierischii
6. Sepals 4.5–8.5 mm; petals 8–15 mm
S. moorei
7. Leaf blades pedately divided, lobes relatively narrow
S. pedatifida
7. Leaf blades usually 3–5-lobed, sometimes pedately divided with relatively broad lobes
→ 8
8. Leaf blades 3–5-lobed or pedately divided, green to gray-green, stellate-canescent; bractlets deciduous.
S. coccinea
8. Leaf blades 3–5-lobed, yellow-green, sparsely stellate-canescent, bractlets persistent.
S. psoraloides
9. Inflorescences usually racemose, sometimes paniculate
→ 10
9. Inflorescences paniculate
→ 15
10. Leaf blades 3-lobed to pedately divided; inflorescences racemose.
S. digitata
10. Leaf blades unlobed, 3-lobed, or pedately divided; inflorescences racemose or paniculate
→ 11
11. Stems decumbent; leaf blades abaxially white to silvery, adaxially green; coastal Texas.
S. lindheimeri
11. Stems usually ascending to erect, sometimes decumbent; leaf blades green, yellow-green, or gray-green; w Texas and sw United States
→ 12
12. Leaf blades ovate to deltate, unlobed or 3-lobed
→ 13
12. Leaf blades oblong-ovate to ovate-lanceolate and unlobed, or widely ovate or triangular to orbiculate and pedately divided
→ 14
13. Stems 5–10 dm; widely distributed.
S. ambigua
13. Stems 2–2.5 dm; restricted to Beaver and Millard counties, Utah, or Nye County, Nevada.
S. caespitosa
14. Leaf blades unlobed; sepals forming beak in bud.
S. hastulata
14. Leaf blades usually pedately divided (all but proximalmost); sepals not forming beak in bud.
S. wrightii
15. Inflorescences open
→ 16
16. Leaf blades unlobed or weakly 3-lobed; stems yellow- to yellow-green canescent or gray- to white-canescent; anthers yellow, gray, or purple; petals red-orange, apricot, lavender, pink, or white.
S. ambigua
16. Leaf blades 3-lobed or 3- or 5-parted; stems gray- to white-tomentose; anthers usually purple; petals red-orange.
S. laxa
17. Leaf blades linear to lanceolate or broadly trullate, unlobed to hastately lobed to angulate; inflorescences usually with leaves throughout.
S. angustifolia
18. Leaf blades unlobed, elongate-deltate, oblong-ovate, ovate-lanceolate, lanceolate, triangular, ovate, or deltate, or weakly 3–5-lobed
→ 19
18. Leaf blades usually strongly 3–5-lobed or pedately divided
→ 22
19. Stems yellow to yellow-green, rubbery; leaf blades deltate or elongate-deltate, sometimes lobed, center lobe sometimes elongate, lobes relatively broad; sepal tips not forming beak in bud.
S. incana
20. Leaf blades oblong-ovate to ovate-lanceolate; stems 1–3(–5) dm; sepal tips forming beak in bud.
S. hastulata
20. Leaf blades ovate to triangular, unlobed or weakly lobed; stems 1.5–10 dm; sepal tips not forming beak in bud
→ 21
21. Stems green to gray-green canescent; leaf margins coarsely toothed; primarily Nevada, Utah, and north.
S. munroana
21. Stems white- to yellow-canescent; leaf margins entire or crenate to serrate; primarily Nevada, Utah, and south.
S. parvifolia
22. Leaf blades 3(–5)-lobed; stems (2–)4–30 dm
→ 23
22. Leaf blades usually deeply lobed or pedately divided, sometimes highly dissected; stems 1–10 dm
→ 26
23. Stems (2–)4–7(–10) dm, usually sparsely canescent or hirsute; petals red-orange.
S. fendleri
23. Stems 10–30 dm, glabrous or hairy; petals red-orange, red, pink, white, lavender, purple, or rose-purple
→ 24
24. Leaf blades ovate-triangular to lanceolate, 3-lobed; inflorescence tip not leafy; stem glabrous or coarse-canescent.
S. emoryi
24. Leaf blades deltate, lanceolate, ovate, triangular, or cordate, subhastate to 3-lobed; inflorescence tip leafy; stem indument soft
→ 25
25. Mericarps 12–14, 3.5–5.5 mm.
S. polychroma
25. Mericarps 10, usually 3 mm.
S. procera
26. Stems (2–)4–10 dm
→ 27
26. Stems 1–3(–6) dm
→ 28
27. Leaf blade surfaces hirsute, sparsely pubescent, or densely soft-pubescent, lobed, appearing green to gray-green.
S. fendleri
27. Leaf blade surfaces stellate-pubescent, commonly lobed or pedately divided, appearing yellow-green to gray-green to green.
S. grossulariifolia
28. Stems green to gray-green; bractlets and leaf margins not red, reddish, or purple, usually green or tan.
S. hastulata
28. Stems gray-green to purple (red-purple basally in S. fumariensis); bractlets and sometimes leaf margins red, reddish, or purple
→ 29
29. Leaf blades densely stellate-pubescent; anthers yellow; mericarps 10–14, 3–4 mm.
S. fumariensis
29. Leaf blades sparsely stellate-pubescent; anthers usually red to purple, sometimes yellow; mericarps 13, 4.5–6 mm.
S. rusbyi
Source FNA vol. 6, p. 357. Author: John La Duke. FNA vol. 6, p. 368.
Parent taxa Malvaceae > subfam. Malvoideae Malvaceae > subfam. Malvoideae > Sphaeralcea
Sibling taxa
S. ambigua, S. angustifolia, S. caespitosa, S. coccinea, S. coulteri, S. digitata, S. emoryi, S. fendleri, S. fumariensis, S. gierischii, S. grossulariifolia, S. hastulata, S. incana, S. laxa, S. leptophylla, S. lindheimeri, S. moorei, S. munroana, S. orcuttii, S. parvifolia, S. pedatifida, S. procera, S. psoraloides, S. rusbyi, S. wrightii
Subordinate taxa
S. ambigua, S. angustifolia, S. caespitosa, S. coccinea, S. coulteri, S. digitata, S. emoryi, S. fendleri, S. fumariensis, S. gierischii, S. grossulariifolia, S. hastulata, S. incana, S. laxa, S. leptophylla, S. lindheimeri, S. moorei, S. munroana, S. orcuttii, S. parvifolia, S. pedatifida, S. polychroma, S. procera, S. psoraloides, S. rusbyi, S. wrightii
Name authority A. St.-Hilaire: in A. St.-Hilaire et al., Fl. Bras. Merid. 1(fol.): 163. (1825) La Duke: SouthW. Naturalist 30: 433, fig. 1. (1985)
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