Silene acaulis |
Silene nivea |
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moss campion, silène acaule |
evening campion, snowy campion |
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Habit | Plants perennial, mat- or cushion-forming, subglabrous; taproot stout; caudex much-branched, becoming woody. | Plants perennial, rhizomatous; rhizome elongate. |
Stems | erect, simple to sparingly branched, leafy, 20–70 cm, glabrous to puberulent, especially distally. |
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Flowering stems | erect, leafy proximally, 3–6(–15) cm, old leaves persistent at base. |
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Leaves | mostly basal, densely crowded and imbricate, sessile; blade 1(–3)-veined, linear-subulate to lanceolate, 0.4–1(–1.5) cm × 0.8–1.5(–2) mm, margins cartilaginous, often ciliolate especially proximally, apex acute, glabrous to scabrous. |
2 per node, sessile or short-petiolate, largest near mid-stem region, reduced and withering proximally, blade elliptic-lanceolate, base cuneate or rounded, apex gradually acuminate and acute, glabrous to puberulent. |
Inflorescences | solitary flowers. |
cymose, (1–)3–5(–12)-flowered, open, leafy. |
Pedicels | 2–40 mm. |
1/2–21/2 times calyx, apex often becoming deflexed, glabrous to hirsute. |
Flowers | bisexual or unisexual, all plants having both staminate and pistillate flowers, others having only pistillate flowers, subsessile or borne singly on peduncle; calyx 10-veined, lateral veins absent, tubular to campanulate, (5–)7–10 mm, herbaceous, margins often purple tinged, dentate, sometimes ciliate, ± scarious, glabrous, lobes lanceolate to ovate, 1–2 mm; petals bright pink, rarely white, limb unlobed to shallowly 2-fid, 2.5–3.5 mm, base tapered into claw, auricles and appendages poorly developed; stamens exserted in staminate flowers, not so or aborted in pistillate flowers; styles 3. |
calyx green, obscurely 10-veined, broadly tubular to campanulate, ± constricted at base around carpophore with broad umbilicate base, becoming broadly clavate in fruit, 14–17 × 5–9 mm, herbaceous, glabrous or hirsute, veins green, without pale commissures, lobes triangular, 2–3 mm; corolla white, clawed, claw equaling calyx, expanded distally into 2-lobed limb, limb oblong, 6–7 mm, appendages oblong, 1–1.6 mm, margins ± entire; stamens short-exserted; stigmas 3, short-exserted. |
Capsules | 3-locular, cylindric, equaling or to 2 times calyx, opening by 6 recurved teeth; carpophore ca. 1 mm. |
globose, equaling calyx, opening by 3 broad teeth that sometimes split to form 6; carpophore 5–6 mm. |
Seeds | light brown, reniform, 0.8–1(–1.2) mm broad, dull, shallowly rugose. |
dark brown to black, with grayish bloom, broadly reniform, not winged, 0.7–1 mm, sides with concentric crescents of low tubercles, larger and deeper on outer margins. |
2n | = 24. |
= 48. |
Silene acaulis |
Silene nivea |
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Phenology | Flowering early summer. | Flowering late spring–summer. |
Habitat | Arctic and alpine tundra, gravelly, often wet places, rocky ledges | Alluvial woodlands |
Elevation | 0-4200 m (0-13800 ft) | 0-400 m (0-1300 ft) |
Distribution |
AK; AZ; CO; ID; ME; MT; NH; NM; NV; NY; OR; UT; WA; WY; AB; BC; NL; NS; NT; NU; ON; QC; SK; YT; Greenland; Europe; Asia (Russian Far East)
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DC; IA; IL; IN; MA; MD; ME; MN; MO; OH; PA; VA; WI; WV
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Discussion | Silene acaulis is a variable species, and most workers have recognized infraspecific taxa in North America: subsp. acaulis (subsp. exscapa and subsp. arctica), which is predominantly arctic; and subsp. subacaulescens, which extends down the Rocky Mountains from Alaska to Arizona and New Mexico. In subsp. acaulis, the leaves are flat and short and the flowers are subsessile and smaller in size. Subspecies subacaulescens is typically a larger, less-compact plant with longer, narrower leaves and larger, pedunculate flowers. However, in many populations, these two variants are poorly differentiated, and in others both occur together, connected by intermediates. Silene acaulis is widely distributed in arctic and alpine Europe. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The green, obscurely veined, umbilicate calyx with its broad base constricted around the carpophore is unique among the North American members of the genus. Silene nivea is occasionally weedy. It was introduced near Québec City (ca. 1969) but did not persist, and probably is not native also in Maine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 173. | FNA vol. 5, p. 194. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Cucubalus acaulis, S. acaulis subsp. arctica, S. acaulis var. exscapa, S. acaulis subsp. subacaulescens, S. exscapa, Xamilensis acaulis | Cucubalus niveus, S. alba |
Name authority | (Linnaeus) Jacquin: Enum. Stirp. Vindob., 78, 242. (1762) | (Nuttall) Muhlenberg ex Otth: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 1: 377. (1824) |
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