Sidalcea nelsoniana |
Sidalcea hirtipes |
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Nelson's checker-mallow, Nelson's checkerbloom |
bluff mallow, bristly-stem checker-mallow, bristly-stem checkerbloom, hairy-stem checkermallow |
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Habit | Herbs, perennial, 0.4–1 m, often glaucous, with thick, woody taproot and lateral rhizomes to 500 mm. | Herbs, perennial, usually in colonies, 0.7–1.3(–1.8) m, not glaucous, with thick, rather woody taproot and coarse, elongate (cordlike) rhizomes 20–100 × 5 mm. |
Stems | clustered, erect, base decumbent-ascending, solid or ± hollow in age, usually ± glaucous distally, glabrous or sparsely hirtellous, hairs short, appressed, simple. |
several, scattered, erect, arising from rhizome apices, usually slightly hollow, densely, harshly bristly-hirsute, hairs stiff, pustular, simple, forked, or stellate, often 2–2.5 mm. |
Leaves | basal and cauline; stipules sometimes deciduous, lanceolate to ovate, (4–)7–9(–14) × (1.2–)3–4 mm; petioles of proximal leaves 18–37 cm, 3–5 times as long as blades, distal 5–20 cm, to 1/2–1 times as long as blades; blades: basal reniform to rounded, unlobed and marginally deeply crenate or very shallowly palmately 5–7-lobed, 6–20 × 6–20 cm, base cordate, apex rounded; cauline deeply 5–9-lobed, lobes linear-elliptic to oblong-elliptic, margins usually entire, sometimes toothed, surfaces finely, sparsely hairy, hairs mostly simple. |
basal and cauline; stipules linear-subulate, 6–8 × 1–1.5 mm; petioles of proximal leaves 20–30 cm, 3 times as long as blades, reduced distally to 1/2 times as long as blades; blades: basal and proximalmost orbiculate to reniform, shallowly 5–9-lobed, 10–15 × 10–15 cm, thick, base narrowly cordate, lobe margins coarsely crenate-dentate, apex rounded, surfaces coarsely hirsute, hairs stiff; distal orbiculate, deeply palmately 5–7-lobed, incised ± to base, lobes sometimes lobed again, base cuneate, apex acute, deeply 2–3-toothed, surfaces long-hirsute or with 2–4-rayed, stellate hairs abaxially. |
Inflorescences | erect, spiciform, proximally open, distally dense, otherwise calyces not conspicuously overlapping except sometimes in bud, branched, 20+-flowered, proximalmost flowers spaced usually to 1 cm apart, not leafy-bracted, elongate, not 1-sided, 10–20 cm; bracts single, purple, mm, hairy; anthers white; stigmas (6 or)7 or 8. |
erect, spiciform to subcapitate, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, proximals usually long-pedunculate, unbranched or branched, 20+-flowered, 10+ flowers usually open on spike at same time, not interrupted, not 1-sided, usually to 8 cm, usually not elongate but sometimes slightly elongated in some populations and to 20 cm in fruit; bracts paired or single, linear, distal undivided, proximal distinct to base, 6 mm, mostly slightly longer than pedicels. |
Pedicels | 1–3(–5) mm; involucellar bractlets absent. |
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Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx often purple tinted, 9–11 mm, to 11–16 mm in fruit, margins ciliate, hairs 1–2 mm, surfaces finely stellate-hairy at base and with coarser, longer, simple and stellate hairs apically; petals usually pale pink to rose-lavender, rarely white, slightly or not pale-veined, (9–)10–21 mm, pistillate often 9–14 mm; staminal column 5–7(–10) mm, hairy; anthers white; stigmas 5–10. |
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Seeds | 1.5 mm. |
2.5–3.5 mm. |
Schizocarps | 4–5 mm diam.; mericarps (6 or)7 or 8, 2 mm, roughened, sparsely glandular-puberulent apically, sides finely reticulate to faintly rugose, sometimes also on back, to nearly smooth, not pitted, mucro 0.5–1 mm. |
7–8 mm diam.; mericarps 5–10, 3.5–4 mm, glabrous or sparsely stellate-puberulent, roughened, prominently reticulate-veined, sides rugose and pitted, back less so, mucro 0.6–0.8 mm. |
2n | = 20. |
= 60. |
Sidalcea nelsoniana |
Sidalcea hirtipes |
|
Phenology | Flowering (May–)Jun–Jul(–Sep). | Flowering (Apr–)May–Jul(–Aug). |
Habitat | Open fields, meadows, fencerows, remnant prairies | Prairie remnants, coastal bluffs, open shrublands, fencerows, meadows, usually mesic, basaltic soil |
Elevation | (40–)100–600(–1300) m ((100–)300–2000(–4300) ft) | 0–200(–1200) m (0–700(–3900) ft) |
Distribution |
OR; WA
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OR; WA
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Discussion | Of conservation concern. Sidalcea nelsoniana is uncommon and persists generally in degraded habitats. It resembles S. campestris and S. virgata, with similar ranges, but is generally less hairy and has a smaller, usually purplish calyx. It has been listed as endangered in Washington and as threatened in Oregon; the number of populations and individuals has greatly declined as a result of land development. It is found in the Willamette Valley area from Multnomah and Washington to Benton and Linn counties in Oregon, and in Cowlitz and Lewis counties in Washington. Sidalcea nelsoniana is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Sidalcea hirtipes is uncommon and known from Clatsop, Lincoln, and Tillamook counties in Oregon and Clark, Lewis, and Wahkiakum counties in Washington. Its elevation and habitat vary, and it seems as much at home on steep coastal cliffs as in more inland, historic prairies and mountain meadows. Populations can appear to be large because of the long-rhizomatous and clonal nature of the plants; they are few and local. It is threatened by grazing, loss of habitat, fire suppression, road construction and maintenance, and changes in hydrology. It is a candidate for listing in Oregon and has been listed as endangered in Washington. Sidalcea hirtipes is characterized by its coarse indument of bristle hairs, its generally compact spikelike inflorescences, its relatively few, large, erect, hirsute leaves, and, especially, its extensive, coarse rhizomes. The inflorescences in some populations are elongated in fruit; its range, hirsute indument, and thick leaves along with coarse rhizomes help to distinguish it from other species. Stem internode length varies depending on habitat, as in many other Sidalcea. Molecular data suggest a relationship among S. hirtipes and S. asprella, S. celata, and S. gigantea (K. Andreasen and B. G. Baldwin 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 346. | FNA vol. 6, p. 339. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Piper: Proc. Biol. Soc. Wash. 32: 41. (1919) | C. L. Hitchcock: Perenn. Sp. Sidalcea, 42. (1957) |
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