Sidalcea nelsoniana |
Sidalcea cusickii |
|
---|---|---|
Nelson's checker-mallow, Nelson's checkerbloom |
Cusick's checkerbloom, Cusick's checkermallow, Cusick's sidalcea |
|
Habit | Herbs, perennial, 0.4–1 m, often glaucous, with thick, woody taproot and lateral rhizomes to 500 mm. | Herbs, perennial, 0.4–1.8 m, not glaucous, with thick taproot and freely-rooting rhizomes usually 3–10 mm diam. |
Stems | clustered, erect, base decumbent-ascending, solid or ± hollow in age, usually ± glaucous distally, glabrous or sparsely hirtellous, hairs short, appressed, simple. |
usually several, clustered, erect, often purplish, often thick and hollow proximally, usually proximally ± scabrous, hairs stellate and distally stellate-hairy, sometimes glabrous. |
Leaves | basal and cauline; stipules sometimes deciduous, lanceolate to ovate, (4–)7–9(–14) × (1.2–)3–4 mm; petioles of proximal leaves 18–37 cm, 3–5 times as long as blades, distal 5–20 cm, to 1/2–1 times as long as blades; blades: basal reniform to rounded, unlobed and marginally deeply crenate or very shallowly palmately 5–7-lobed, 6–20 × 6–20 cm, base cordate, apex rounded; cauline deeply 5–9-lobed, lobes linear-elliptic to oblong-elliptic, margins usually entire, sometimes toothed, surfaces finely, sparsely hairy, hairs mostly simple. |
mostly cauline; stipules persistent or deciduous, lanceolate, 6–8 × 1–1.5 mm; proximal petioles 25–30 cm, 4–5 times as long as blade, reduced to 7–13 cm on midstem leaves and to 2–3 cm on distalmost leaves, these to 1/2 times as long as blades; basal blades wide-ovate to orbiculate, shallowly to deeply, palmately 5–7(–9)-lobed, 6–13 × 6–13 cm, lobe margins toothed, teeth rounded, sometimes Ribes-like, base cordate, apex rounded, surfaces: abaxial densely stellate-pubescent, adaxial glabrous or sparsely stellate-pubescent on veins; cauline blades round, divided nearly to base, 5–9-lobed, lobes again deeply incised to subentire, 10–20 × 10–20 cm, distalmost 5–7-cleft to base. |
Inflorescences | erect, spiciform, proximally open, distally dense, otherwise calyces not conspicuously overlapping except sometimes in bud, branched, 20+-flowered, proximalmost flowers spaced usually to 1 cm apart, not leafy-bracted, elongate, not 1-sided, 10–20 cm; bracts single, purple, mm, hairy; anthers white; stigmas (6 or)7 or 8. |
erect, spiciform, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually 20–30-branched per primary stem, 20+-flowered, elongate, not 1-sided, most flowers usually open at same time, branches relatively short, each spike unit 6 cm, longer in fruit; bracts ovate-lanceolate to subulate, undivided, 1–6(–10) mm, subequal or longer than pedicels, much shorter than calyx. |
Pedicels | 1–2(–5) mm; involucellar bractlets absent. |
|
Flowers | bisexual or unisexual and pistillate, pistillate plants more frequent than bisexual ones, some flowers may be staminate, plants gynodioecious; calyx urceolate with swollen base especially in young fruit, pistillate 6–8 mm, bisexual 6–10 mm, strongly reticulate-veined, usually finely stellate-puberulent, sometimes glabrate, lobes usually purple tinted; petals 7 or 8, pale pink, pinkish rose, or rose-purple, usually neither pale-veined nor white at base, pistillate 8–12(–14) mm, staminate or bisexual (10–)11–19(–23) mm; staminal column 5–6 mm, hairy; anthers white; stigmas 7 or 8. |
|
Seeds | 1.5 mm. |
2 mm. |
Schizocarps | 4–5 mm diam.; mericarps (6 or)7 or 8, 2 mm, roughened, sparsely glandular-puberulent apically, sides finely reticulate to faintly rugose, sometimes also on back, to nearly smooth, not pitted, mucro 0.5–1 mm. |
6 mm diam.; mericarps 7 or 8, 3 mm, apical margins ± rounded, sides smooth or slightly reticulate-veined, not pitted, back essentially smooth, very sparsely glandular-puberulent near tip, mucro 0.5–1 mm. |
2n | = 20. |
= 20. |
Sidalcea nelsoniana |
Sidalcea cusickii |
|
Phenology | Flowering (May–)Jun–Jul(–Sep). | Flowering Jun–Aug. |
Habitat | Open fields, meadows, fencerows, remnant prairies | Moist to wet, mostly black, adobe soil, lowland and mountain meadows, often with Juncus and Camassia |
Elevation | (40–)100–600(–1300) m ((100–)300–2000(–4300) ft) | 100–200(–500), 1000–1400 m (300–700(–1600), 3300–4600 ft) |
Distribution |
OR; WA
|
OR
|
Discussion | Of conservation concern. Sidalcea nelsoniana is uncommon and persists generally in degraded habitats. It resembles S. campestris and S. virgata, with similar ranges, but is generally less hairy and has a smaller, usually purplish calyx. It has been listed as endangered in Washington and as threatened in Oregon; the number of populations and individuals has greatly declined as a result of land development. It is found in the Willamette Valley area from Multnomah and Washington to Benton and Linn counties in Oregon, and in Cowlitz and Lewis counties in Washington. Sidalcea nelsoniana is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea cusickii is showy and distinctive. It has been confused with S. campestris, S. hendersonii, S. nelsoniana, S. oregana (subsp. spicata), S. setosa, and S. virgata. Molecular data support a close relationship with S. hendersonii and S. virgata (K. Andreasen and B. G. Baldwin 2003b). Hitchcock distinguished subsp. cusickii and subsp. purpurea, the latter with purple, ciliate calyx lobes and stem proximally glabrous and thought to be more closely related to S. hendersonii and S. nelsoniana. The two variations overlap in range and characteristics and are not recognized here. There appear to be two groups of populations at different elevation ranges with little overlap. Overall, S. cusickii can be distinguished by its range and by its spikelike, many-flowered, thick, compounded racemes in which essentially all of the flowers are open at the same time. It is also unusual in the preponderance of pistillate-flowered individuals in a given population and the presence of what appear to be truly staminate, rather than bisexual, flowers on some individuals. It is found in the Willamette and Umpqua valley region in Douglas, Josephine, and Lane counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 346. | FNA vol. 6, p. 330. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. cusickii subsp. purpurea, S. oregana var. cusickii | |
Name authority | Piper: Proc. Biol. Soc. Wash. 32: 41. (1919) | Piper: Proc. Biol. Soc. Wash. 29: 99. (1916) |
Web links |