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chaparral checkerbloom, Hickman's checkerbloom

Del Norte checkerbloom, dwarf checkerbloom

Habit Herbs, perennial, 0.1–0.8 m, not glaucous, with thick, woody taproot or caudex, without rhizomes. Herbs, perennial, 0.2–0.6(–0.8) m, sometimes ± glaucous, with woody taproot or caudex and rhizomes, freely rooting, 20–30 cm × 2–4 mm, often mat-forming, forming clones often 1–8 m2.
Stems

several to many (ca. 3–20+), clustered, erect to ascending, branched or unbranched, solid, usually densely stellate-canescent.

scattered, erect to ascending, base decumbent to erect, usually rooting freely, solid, proximally hairy or glabrate, hairs soft, simple and stellate, distally brittle, easily broken, glabrous-glaucous.

Leaves

cauline, evenly arrayed on stem, usually similar in size, shape;

stipules linear-lanceolate to ovate, 2–9 × 1–3 mm, widest above base, width sometimes exceeding stem diam.;

petiole 0.6–3(–9) cm, 1/2–3 times as long as blade, apex often with pulvinus;

blade orbiculate or reniform to flabelliform, unlobed and margins coarsely crenate to shallowly or deeply lobed, 1–7 × 1–7 cm, usually wider than long, base truncate or cordate, apex rounded, surfaces stellate-hairy.

mostly basal or cauline 3 or 4 on proximal 1/3 of stem, much reduced;

stipules wide-lanceolate to ovate, 3–4 × 1–1.5 mm;

petioles of proximal leaves 6–12 cm, 2–4 times as long as blades in basal leaves, those of cauline leaves greatly reduced to 1/2 times or as long as blades;

blade rounded to reniform, palmately (3–)5–7-lobed, usually (1–)2–5(–10) × (1–)2–5(–10) cm, apex rounded, surfaces: abaxial harshly stellate-hairy, adaxial usually simple-haired, basal blades shallowly incised, lobes with 3 deep crenations;

cauline blades usually deeply 3–5(–7)-lobed nearly to base, lobe margins dentate or entire.

Inflorescences

erect, infrequently ascending, usually spiciform, dense or open, calyces overlapping or not, branched or unbranched, 2–20+-flowered, proximal flowers scattered, usually more congested distally, not notably elongate in flower, not 1-sided, (1.5–)3–25 cm, usually longer in fruit;

bracts linear to ovate-lanceolate or oblong, undivided, 2-fid, or divided, 2–8(–12) mm, not involucrelike, distal entire to 2-fid, stipulelike, proximalmost not involucrelike, divided to base, much shorter than to nearly equaling calyx.

ascending, open, calyces not conspicuously overlapping except sometimes in bud, unbranched or few-branched, loosely (3–)5–10(–20)-flowered, not greatly elongated, 1-sided, 10–20 cm;

bracts narrowly elliptic, shallowly 2-fid, proximal bracts divided to base and often with leaf remnant between, 3–4 mm, usually shorter than to equaling pedicels.

Pedicels

1–4(–5) mm;

involucellar bractlets (2 or)3, 2–10 mm, shorter to slightly longer than calyx.

3–4(–10) mm;

involucellar bractlets absent.

Flowers

usually bisexual, infrequently unisexual and pistillate;

calyx 4–12 mm, densely to sparsely stellate-puberulent to long-bristly;

petals usually pale pink to pink-lavender, rarely white, veins not conspicuously whitened, 5–17 mm;

staminal column 4–7 mm, hairy;

anthers white to pale pinkish or pale yellow;

stigmas (4–)6 or 7(–10).

bisexual or unisexual and pistillate, plants gynodioecious;

calyx 7–10 mm, slightly enlarged in fruit, uniformly, coarsely stellate-puberulent (some rays sometimes longer than others);

petals dark pink, pale-veined at least when dry, pistillate 10–15(–20) mm, bisexual 20–25(–33) mm;

staminal column 4–5 mm, sparsely hairy;

anthers white to pale pink;

stigmas 6 or 7(or 8).

Seeds

1–2 mm.

2 mm.

Schizocarps

4–7 mm diam.;

mericarps usually (4–)6 or 7(–10), (1.5–)2–2.5 mm, glabrous, sides usually smooth, thin, margins and back usually lightly reticulate-veined, transversely corrugated, back usually with medial, raised line, not pitted, mucro absent.

6–7 mm diam.;

mericarps 6 or 7(or 8), 3–3.5 mm, back not ribbed, roughened, sides and back strongly reticulate-veined-rugose and pitted (honeycomblike), top minutely glandular-puberulent, mucro 0.8–1 mm.

2n

= 40, 60.

Sidalcea hickmanii

Sidalcea elegans

Phenology Flowering (May–)Jun–Jul(–Sep).
Habitat Open, dry woodlands, usually on serpentine
Elevation 100–200(–900) m (300–700(–3000) ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Sidalcea hickmanii is found in isolated populations from southern California to southwestern Oregon and appears to have a relict distribution. K. Andreasen and B. G. Baldwin (2001, 2003) suggested that it is basal within Sidalcea. It is distinctive in having three (normally two in subsp. petraea) involucellar bractlets attached to the calyx, no mucro on the mericarps, and leaves that are almost the same size and shape throughout the stem. Each subspecies apparently represents a distinct relictual colony; the sexuality of these is not well known because of the paucity of specimens. As in many sidalceas, this species in particular appears to be fire-dependent.

Subspecies 7 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Sidalcea elegans has been included within S. malviflora; it is easily distinguished by its relatively long, slender, shallow rhizomes; open, one-sided inflorescences; and thin, brittle stems. It resembles S. glaucescens in its leaves and inflorescence and is easily distinguished by its rhizomes and soft, simple hairs at the stem base. It has been confused with S. asprella, from which it is distinguished by its simple, flexible hairs at the stem base and by its more developed and elongated rhizomes and variable leaves. Sidalcea elegans occurs in northwestern California and southwestern Oregon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades usually lobed, incised ± to base; California
→ 2
1. Leaf blades unlobed, incised to 1/2 length; California, Oregon
→ 3
2. Bracts (7–)10–12 mm, equaling or shorter than calyx; c San Luis Obispo County.
subsp. anomala
2. Bracts 5.5–7 mm, shorter than calyx; Napa County.
subsp. napensis
3. Involucellar bractlets 2(–3); petals white to pale pink; flowers bisexual or pistillate; sw Oregon.
subsp. petraea
3. Involucellar bractlets 3; petals pink, pale pink, pinkish lavender, or pale lavender; flowers bisexual; California
→ 4
4. Bracts broadly lanceolate, 5–7(–10) × 2.5–4 mm, slightly shorter than calyx; involucellar bractlets equaling or slightly shorter than calyx; leaf blades: distal unlobed or incised to 1/4 length; Santa Barbara and San Bernardino counties.
subsp. parishii
5. Plants 0.4–0.8 m; stems brick red, greenish, or grayish; calyces stellate-puberulent, hairs longest at margins; involucellar bractlets 2–7 mm; largest leaf blades deeply cordate, 2.5–7 cm wide; inflorescences dense; Monterey County.
subsp. hickmanii
6. Bracts of distal flowers 1, cupped; leaf blades 0.6–1.5 × 0.7–2.2 cm; stems distally stellate-hairy, hairs appressed, 0.2–0.5 mm; plants 0.1–0.4 m; inflorescences not spiciform, to 10-flowered; calyces 4–5.5 mm; n Lake County.
subsp. pillsburiensis
6. Bracts of distal flowers usually 2, flat or cupped; leaf blades (1–)2–4 × (1–)2.7 cm; stems distally hairy, hairs tufted, 0.5–1.2 mm; plants (0.2–)0.3(–0.4) m; inflorescences spiciform in age, 10+-flowered; calyces 6–7 mm; Marin County.
subsp. viridis
Source FNA vol. 6, p. 334. FNA vol. 6, p. 331.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. hickmanii subsp. anomala, S. hickmanii subsp. hickmanii, S. hickmanii subsp. napensis, S. hickmanii subsp. parishii, S. hickmanii subsp. petraea, S. hickmanii subsp. pillsburiensis, S. hickmanii subsp. viridis
Synonyms S. malviflora subsp. elegans
Name authority Greene: Pittonia 1: 139. (1887) Greene: Cybele Columb. 1: 35. (1914)
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