Sidalcea hickmanii |
Sidalcea campestris |
|||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
chaparral checkerbloom, Hickman's checkerbloom |
meadow checker-mallow, meadow checkerbloom, meadow sidalcea |
|||||||||||||||||||||
Habit | Herbs, perennial, 0.1–0.8 m, not glaucous, with thick, woody taproot or caudex, without rhizomes. | Herbs, perennial, 0.5–2 m, sometimes glaucous in age, with thick, rather woody taproot and short rhizomes usually to 10 cm × 10 mm. | ||||||||||||||||||||
Stems | several to many (ca. 3–20+), clustered, erect to ascending, branched or unbranched, solid, usually densely stellate-canescent. |
single or clustered, erect, base often decumbent-ascending, solid (proximally hollow on older stems), proximally densely bristly-hirsute, hairs simple or forked, 0.5–1 mm, sometimes mixed with minute, stellate hairs, sometimes glabrous and glaucous in age, hairs usually more appressed, simple, stellate, or sparse distally. |
||||||||||||||||||||
Leaves | cauline, evenly arrayed on stem, usually similar in size, shape; stipules linear-lanceolate to ovate, 2–9 × 1–3 mm, widest above base, width sometimes exceeding stem diam.; petiole 0.6–3(–9) cm, 1/2–3 times as long as blade, apex often with pulvinus; blade orbiculate or reniform to flabelliform, unlobed and margins coarsely crenate to shallowly or deeply lobed, 1–7 × 1–7 cm, usually wider than long, base truncate or cordate, apex rounded, surfaces stellate-hairy. |
mostly cauline; stipules lanceolate, 4–11 × 1–1.5 mm; petioles of proximal leaves 15–20 cm, 2–3 times as long as blades, distal reduced to 0.5–10 cm, 1/2 times to as long as blades; blade 10–15 × 10–15 cm, smaller distally, surfaces scabrid-hairy, hairs dense, simple or stellate, stiff, stellate hairs mostly on adaxial surfaces, proximal blades orbiculate, shallowly to deeply palmately 7–9-lobed, 5–15 × 5–15 cm, base cordate, margins coarsely crenate-serrate, lobes apically 2–5-toothed; distal cauline leaves variable, mid blades usually palmately divided nearly to base into 5–7 cuneate lobes, 15 × 15 cm, lobes deeply cut to laciniately dissected, distal blades divided into (3–)5–7 linear, marginally subentire segments, surfaces glabrescent or with few hairs on abaxial surface veins, ciliate. |
||||||||||||||||||||
Inflorescences | erect, infrequently ascending, usually spiciform, dense or open, calyces overlapping or not, branched or unbranched, 2–20+-flowered, proximal flowers scattered, usually more congested distally, not notably elongate in flower, not 1-sided, (1.5–)3–25 cm, usually longer in fruit; bracts linear to ovate-lanceolate or oblong, undivided, 2-fid, or divided, 2–8(–12) mm, not involucrelike, distal entire to 2-fid, stipulelike, proximalmost not involucrelike, divided to base, much shorter than to nearly equaling calyx. |
erect, open, spiciform, calyces not conspicuously overlapping except sometimes in bud, often branched from distal leaves, 15+-flowered, elongate, not 1-sided, 10–35 cm, proximal flowers spaced several cm apart, not leafy-bracted; bracts linear, distal undivided or 2-fid, proximal separate to base, 4–10 mm, usually equaling or longer than pedicels. |
||||||||||||||||||||
Pedicels | 1–4(–5) mm; involucellar bractlets (2 or)3, 2–10 mm, shorter to slightly longer than calyx. |
3–6(–20) mm; involucellar bractlets absent. |
||||||||||||||||||||
Flowers | usually bisexual, infrequently unisexual and pistillate; calyx 4–12 mm, densely to sparsely stellate-puberulent to long-bristly; petals usually pale pink to pink-lavender, rarely white, veins not conspicuously whitened, 5–17 mm; staminal column 4–7 mm, hairy; anthers white to pale pinkish or pale yellow; stigmas (4–)6 or 7(–10). |
bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–9 mm, pistillate 5–7 mm, bisexual 6–9 mm, 8–10 mm in fruit, uniformly, densely stellate-hairy or with coarser, longer, stellate hairs to 2 mm; petals usually not overlapping, nearly white to pale pink or pale lavender, pistillate 9–12 mm, bisexual 13–25 mm; staminal column 5–7 mm, hairy; anthers white to pale pink; stigmas (6 or)7 or 8. |
||||||||||||||||||||
Seeds | 1–2 mm. |
2.5 mm. |
||||||||||||||||||||
Schizocarps | 4–7 mm diam.; mericarps usually (4–)6 or 7(–10), (1.5–)2–2.5 mm, glabrous, sides usually smooth, thin, margins and back usually lightly reticulate-veined, transversely corrugated, back usually with medial, raised line, not pitted, mucro absent. |
7–8 mm diam.; mericarps (6 or)7 or 8, 3.5 mm, roughened, sides prominently reticulate-rugose and pitted, back less so and glandular-puberulent, mucro 0.5–1 mm. |
||||||||||||||||||||
2n | = 60. |
|||||||||||||||||||||
Sidalcea hickmanii |
Sidalcea campestris |
|||||||||||||||||||||
Phenology | Flowering May–Aug. | |||||||||||||||||||||
Habitat | Open shrublands, meadows, hedgerows, prairies | |||||||||||||||||||||
Elevation | 40–200 m (100–700 ft) | |||||||||||||||||||||
Distribution |
CA; OR
|
OR
|
||||||||||||||||||||
Discussion | Sidalcea hickmanii is found in isolated populations from southern California to southwestern Oregon and appears to have a relict distribution. K. Andreasen and B. G. Baldwin (2001, 2003) suggested that it is basal within Sidalcea. It is distinctive in having three (normally two in subsp. petraea) involucellar bractlets attached to the calyx, no mucro on the mericarps, and leaves that are almost the same size and shape throughout the stem. Each subspecies apparently represents a distinct relictual colony; the sexuality of these is not well known because of the paucity of specimens. As in many sidalceas, this species in particular appears to be fire-dependent. Subspecies 7 (7 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea campestris is one of the taller species of Sidalcea and can be distinguished also by its range, hirsute indument, long inflorescences with spaced, long-stalked flowers, narrow very pale petals, and deeply incised pinnatifid stem leaves. It has been confused with S. hirtipes, S. nelsoniana, and S. oregana; it differs especially in characters of rhizomes, inflorescences, and leaves. It is a candidate for listing as threatened or endangered in Oregon. Specimens from British Columbia and Washington identified as S. campestris are either S. campestris escaped from cultivation or S. hendersonii. It is known as a native only from the Willamette Valley area (Multnomah and Washington to Benton and Linn counties). Apparently, it was introduced near Seattle, Washington; it may not persist there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||
Key |
|
|||||||||||||||||||||
Source | FNA vol. 6, p. 334. | FNA vol. 6, p. 328. | ||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||
Synonyms | S. asplenifolia, S. sylvestris | |||||||||||||||||||||
Name authority | Greene: Pittonia 1: 139. (1887) | Greene: Bull. Calif. Acad. Sci. 1: 76. (1885) | ||||||||||||||||||||
Web links |