Sidalcea glaucescens
Malvaceae
glaucous checker mallow, waxy checker mallow, waxy checkerbloom
mallow family
usually few to many, clustered, sprawling or decumbent to ascending, rarely erect, not rooting, solid, glaucous, proximally usually stellate-puberulent, sometimes glabrous, distally glabrous.
basal and cauline, basal leaves 9 or fewer or deciduous;
stipules lanceolate, (2–)3–5(–6) × 0.5–1.5 mm;
petioles of basal and proximal cauline leaves 6–14 cm, 3–4 times as long as blades, reduced distally to 1/2 times to as long as blades;
blade reniform-orbiculate, palmately 5(–7)-lobed, deeply incised, 2–6(–8) × 2–6(–8) cm, glaucous, surfaces glabrous or minutely stellate-puberulent, lobes shallowly dentate, more deeply divided on distal leaves, margins entire, distalmost sometimes linear, unlobed.
alternate, usually spiral, sometimes distichous (Malvoideae), usually petiolate, sometimes subsessile or sessile (Malvoideae), stipulate (usually well developed), simple (compound in Abelmoschus);
blade unlobed or palmately lobed, palmately veined.
ascending, open, calyces not conspicuously overlapping except sometimes in bud, usually unbranched, 3–10(–20)-flowered, elongate, 1-sided, 8–20 cm, axis curved between flowers, sometimes zigzag in appearance;
bracts linear to lanceolate, distinct or connate and 2-fid, 5 mm, proximal divided to base, distal often undivided, shorter than to equaling pedicels.
axillary, terminal, or leaf-opposed.
2–3(–10) mm;
involucellar bractlets absent.
bisexual or unisexual and pistillate, plants gynodioecious;
calyx 5–10 mm, enlarging in fruit, hairy, hairs scattered, minute, stellate and sometimes capitate, glandular;
petals pink to pink-purple, pale-veined at least when dry, pistillate (7–)9–12 mm, bisexual 15–20(–25) mm;
staminal column 4–7 mm, stellate-hairy;
anthers pale yellow or pinkish to white;
stigmas 6–8.
bisexual or unisexual, usually actinomorphic;
involucel (epicalyx) sometimes deciduous (Malvoideae, Sterculioideae), (4–)5(–8), distinct or connate;
petals 4 or 5 (absent in Bombacoideae and Sterculioideae, rarely absent in Grewioideae);
nectaries glandular hairs on adaxial base of sepals, petals, or androgynophores, sometimes absent;
androgynophore present or absent;
stamens [4–]5–100[–1500], usually in antipetalous groups; usually same number as sepals, distinct or connate, sessile or on androgynophore;
ovules (1–)2–many per ovary.
usually capsules, sometimes follicles, schizocarps, berries, or nuts.
2 mm.
cotyledons usually folded, endosperm absent or sparse to copious.
5–7 mm diam.;
mericarps 6–8, 3–3.5 mm, roughened, sides reticulate-veined and deeply pitted, back reticulate-veined and glandular-puberulent, mucro 0.3–1 mm.
= 40.
Sidalcea glaucescens
Malvaceae
Sidalcea glaucescens is a relatively common, generally low-growing plant of relatively high elevations; it occurs from the central and northern Sierra Nevada to the southern Cascade and Klamath ranges and to north of Reno, Nevada. It usually can be distinguished by its highly glaucous, waxy stems and leaves, 3–5-lobed, entire-margined leaves, and basal leaves that wither by the time fruit is mature; additionally, proximal flowers are spaced several centimeters apart and leafy-bracted, and the inflorescence axis is curved between flowers. It has been confused with S. asprella, S. elegans, and S. multifida, to which it appears to be closely related. It can generally be distinguished from S. multifida by its 5(–7)-lobed leaves, the lobes shallowly incised or entire, its nonpersisting, fewer basal leaves, and its more-procumbent habit. Sidalcea elegans and S. virgata in southwestern Oregon also have been confused with S. glaucescens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera ca. 240, species ca. 4350 (52 genera, 250 species in the flora).
Malvaceae comprise taxa traditionally separated among four families: Malvaceae, Bombacaceae Kunth, Sterculiaceae Ventenat, and Tiliaceae Jussieu. Morphological characters distinguishing these previously recognized families are notoriously ambiguous and/or absent. Multiple tribes and genera (for example, Fremontodendron) have been transferred between families as taxonomic boundaries changed throughout history. Molecular phylogenies indicate that only one of the four previously recognized families (Malvaceae in the narrow sense) forms a monophyletic group (C. Bayer et al. 1999; W. S. Alverson et al. 1999) and the monophyly of an expanded familial circumscription, including all four previously accepted families, is well documented (W. S. Judd and S. R. Manchester 1997; Alverson et al. 1998; Bayer et al.). Based on morphological, molecular, and biogeographic data, Malvaceae now include nine subfamilies (Bayer et al.; Alverson et al. 1999), six of which are represented in the flora area; Malvoideae and Bombacoideae form a monophyletic group that is part of a larger clade that includes some
traditional components of Tiliaceae and Sterculiaceae (now Tilioideae and Sterculioideae). A second clade contains Grewioideae and Byttneriodeae. Not included in the flora are members of subfamilies Brownlowioideae Burret, Dombeyoideae Beilschmied, and Helicteroideae Meisner.
Malvaceae range widely in inflorescence structure; all members share a basic repeating bicolor unit (a terminal flower and three bracts or an epicalyx; C. Bayer 1999). Floral nectaries in the family are composed of dense multicellular, glandular hairs on sepals, petals, or androgynophore (S. Vogel 2000). These nectaries provide nectar to a broad range of cells in wood; M. M. Chattaway 1933) are restricted to Malvaceae but are not present in every taxon.
Representatives of Malvaceae are present on every continent except Antarctica; diversity increases in warmer regions. A majority of the genera in Malvoideae and Bombacoideae are native to the New World (P. A. Fryxell 1997); members of Byttnerioideae, Grewioideae, and Sterculioideae are more evenly distributed throughout the Tropics. Tilioideae are restricted to the Northern Hemisphere (C. Bayer and K. Kubitzki 2003).
Malvaceae have an extensive pollen fossil record and a majority of the subfamilies are represented in the Paleocene or Eocene. Tilia (Tilioideae) fossil pollen and leaves are present in western North American temperate forests (where it is now absent) from the mid Eocene (S. R. Manchester 1994; H. W. Meyer and Manchester 1997). North American fossil pollen deposits of Bombacoideae are plentiful in the Cretaceous (W. Krutzsch 1989; B. E. Pfeil et al. 2002 and references therein).
The hairy seed coat of cotton [Gossypium (Malvoideae)] is the most economically valuable product in the family and the historical and evolutionary importance of its domestication is well documented. The seeds of cacao [Theobroma cacao (Sterculioideae)] are the basis of chocolate. Okra [Abelmoschus esculentus (Malvoideae)] is a major vegetable crop in the southeastern United States. Tilia (Tilioideae) trees are planted throughout temperate regions to beautify streets and parks. The marshmallow, Althaea officinalis (Malvoideae), is a perennial herb found in northeastern North America and Europe; the mucilage from its roots was used to make the original marshmallow.
Key to Subfamilies of Malvaceae
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key to Subfamilies of Malvaceae
1. Gynoecium apocarpous; petals absent; epicalyx absent; androgynophore present; flowers functionally unisexual | a. Sterculioideae |
1. Gynoecium syncarpous; petals present or absent; epicalyx present or absent; androgynophore present or absent; flowers usually bisexual | → 2 |
2. Petals absent | b. Bombacoideae |
2. Petals usually present | → 3 |
3. Epicalyx usually absent | → 4 |
3. Epicalyx usually present, rarely absent | → 5 |
4. Androgynophore absent; nectaries on sepals | c. Tilioideae |
4. Androgynophore present or absent; nectaries on petals or androgynophore | d. Grewioideae |
5. Staminodes usually present; anthers 2- or 3-thecate | e. Byttnerioideae |
5. Staminodes absent or relatively small; anthers 1-thecate | f. Malvoideae |
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