Sidalcea cusickii
Malvaceae
Cusick's checkerbloom, Cusick's checkermallow, Cusick's sidalcea
mallow family
usually several, clustered, erect, often purplish, often thick and hollow proximally, usually proximally ± scabrous, hairs stellate and distally stellate-hairy, sometimes glabrous.
mostly cauline;
stipules persistent or deciduous, lanceolate, 6–8 × 1–1.5 mm;
proximal petioles 25–30 cm, 4–5 times as long as blade, reduced to 7–13 cm on midstem leaves and to 2–3 cm on distalmost leaves, these to 1/2 times as long as blades;
basal blades wide-ovate to orbiculate, shallowly to deeply, palmately 5–7(–9)-lobed, 6–13 × 6–13 cm, lobe margins toothed, teeth rounded, sometimes Ribes-like, base cordate, apex rounded, surfaces: abaxial densely stellate-pubescent, adaxial glabrous or sparsely stellate-pubescent on veins;
cauline blades round, divided nearly to base, 5–9-lobed, lobes again deeply incised to subentire, 10–20 × 10–20 cm, distalmost 5–7-cleft to base.
alternate, usually spiral, sometimes distichous (Malvoideae), usually petiolate, sometimes subsessile or sessile (Malvoideae), stipulate (usually well developed), simple (compound in Abelmoschus);
blade unlobed or palmately lobed, palmately veined.
erect, spiciform, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually 20–30-branched per primary stem, 20+-flowered, elongate, not 1-sided, most flowers usually open at same time, branches relatively short, each spike unit 6 cm, longer in fruit;
bracts ovate-lanceolate to subulate, undivided, 1–6(–10) mm, subequal or longer than pedicels, much shorter than calyx.
axillary, terminal, or leaf-opposed.
1–2(–5) mm;
involucellar bractlets absent.
bisexual or unisexual and pistillate, pistillate plants more frequent than bisexual ones, some flowers may be staminate, plants gynodioecious;
calyx urceolate with swollen base especially in young fruit, pistillate 6–8 mm, bisexual 6–10 mm, strongly reticulate-veined, usually finely stellate-puberulent, sometimes glabrate, lobes usually purple tinted;
petals 7 or 8, pale pink, pinkish rose, or rose-purple, usually neither pale-veined nor white at base, pistillate 8–12(–14) mm, staminate or bisexual (10–)11–19(–23) mm;
staminal column 5–6 mm, hairy;
anthers white;
stigmas 7 or 8.
bisexual or unisexual, usually actinomorphic;
involucel (epicalyx) sometimes deciduous (Malvoideae, Sterculioideae), (4–)5(–8), distinct or connate;
petals 4 or 5 (absent in Bombacoideae and Sterculioideae, rarely absent in Grewioideae);
nectaries glandular hairs on adaxial base of sepals, petals, or androgynophores, sometimes absent;
androgynophore present or absent;
stamens [4–]5–100[–1500], usually in antipetalous groups; usually same number as sepals, distinct or connate, sessile or on androgynophore;
ovules (1–)2–many per ovary.
usually capsules, sometimes follicles, schizocarps, berries, or nuts.
2 mm.
cotyledons usually folded, endosperm absent or sparse to copious.
6 mm diam.;
mericarps 7 or 8, 3 mm, apical margins ± rounded, sides smooth or slightly reticulate-veined, not pitted, back essentially smooth, very sparsely glandular-puberulent near tip, mucro 0.5–1 mm.
= 20.
Sidalcea cusickii
Malvaceae
Sidalcea cusickii is showy and distinctive. It has been confused with S. campestris, S. hendersonii, S. nelsoniana, S. oregana (subsp. spicata), S. setosa, and S. virgata. Molecular data support a close relationship with S. hendersonii and S. virgata (K. Andreasen and B. G. Baldwin 2003b). Hitchcock distinguished subsp. cusickii and subsp. purpurea, the latter with purple, ciliate calyx lobes and stem proximally glabrous and thought to be more closely related to S. hendersonii and S. nelsoniana. The two variations overlap in range and characteristics and are not recognized here. There appear to be two groups of populations at different elevation ranges with little overlap. Overall, S. cusickii can be distinguished by its range and by its spikelike, many-flowered, thick, compounded racemes in which essentially all of the flowers are open at the same time. It is also unusual in the preponderance of pistillate-flowered individuals in a given population and the presence of what appear to be truly staminate, rather than bisexual, flowers on some individuals. It is found in the Willamette and Umpqua valley region in Douglas, Josephine, and Lane counties.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera ca. 240, species ca. 4350 (52 genera, 250 species in the flora).
Malvaceae comprise taxa traditionally separated among four families: Malvaceae, Bombacaceae Kunth, Sterculiaceae Ventenat, and Tiliaceae Jussieu. Morphological characters distinguishing these previously recognized families are notoriously ambiguous and/or absent. Multiple tribes and genera (for example, Fremontodendron) have been transferred between families as taxonomic boundaries changed throughout history. Molecular phylogenies indicate that only one of the four previously recognized families (Malvaceae in the narrow sense) forms a monophyletic group (C. Bayer et al. 1999; W. S. Alverson et al. 1999) and the monophyly of an expanded familial circumscription, including all four previously accepted families, is well documented (W. S. Judd and S. R. Manchester 1997; Alverson et al. 1998; Bayer et al.). Based on morphological, molecular, and biogeographic data, Malvaceae now include nine subfamilies (Bayer et al.; Alverson et al. 1999), six of which are represented in the flora area; Malvoideae and Bombacoideae form a monophyletic group that is part of a larger clade that includes some
traditional components of Tiliaceae and Sterculiaceae (now Tilioideae and Sterculioideae). A second clade contains Grewioideae and Byttneriodeae. Not included in the flora are members of subfamilies Brownlowioideae Burret, Dombeyoideae Beilschmied, and Helicteroideae Meisner.
Malvaceae range widely in inflorescence structure; all members share a basic repeating bicolor unit (a terminal flower and three bracts or an epicalyx; C. Bayer 1999). Floral nectaries in the family are composed of dense multicellular, glandular hairs on sepals, petals, or androgynophore (S. Vogel 2000). These nectaries provide nectar to a broad range of cells in wood; M. M. Chattaway 1933) are restricted to Malvaceae but are not present in every taxon.
Representatives of Malvaceae are present on every continent except Antarctica; diversity increases in warmer regions. A majority of the genera in Malvoideae and Bombacoideae are native to the New World (P. A. Fryxell 1997); members of Byttnerioideae, Grewioideae, and Sterculioideae are more evenly distributed throughout the Tropics. Tilioideae are restricted to the Northern Hemisphere (C. Bayer and K. Kubitzki 2003).
Malvaceae have an extensive pollen fossil record and a majority of the subfamilies are represented in the Paleocene or Eocene. Tilia (Tilioideae) fossil pollen and leaves are present in western North American temperate forests (where it is now absent) from the mid Eocene (S. R. Manchester 1994; H. W. Meyer and Manchester 1997). North American fossil pollen deposits of Bombacoideae are plentiful in the Cretaceous (W. Krutzsch 1989; B. E. Pfeil et al. 2002 and references therein).
The hairy seed coat of cotton [Gossypium (Malvoideae)] is the most economically valuable product in the family and the historical and evolutionary importance of its domestication is well documented. The seeds of cacao [Theobroma cacao (Sterculioideae)] are the basis of chocolate. Okra [Abelmoschus esculentus (Malvoideae)] is a major vegetable crop in the southeastern United States. Tilia (Tilioideae) trees are planted throughout temperate regions to beautify streets and parks. The marshmallow, Althaea officinalis (Malvoideae), is a perennial herb found in northeastern North America and Europe; the mucilage from its roots was used to make the original marshmallow.
Key to Subfamilies of Malvaceae
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key to Subfamilies of Malvaceae
1. Gynoecium apocarpous; petals absent; epicalyx absent; androgynophore present; flowers functionally unisexual | a. Sterculioideae |
1. Gynoecium syncarpous; petals present or absent; epicalyx present or absent; androgynophore present or absent; flowers usually bisexual | → 2 |
2. Petals absent | b. Bombacoideae |
2. Petals usually present | → 3 |
3. Epicalyx usually absent | → 4 |
3. Epicalyx usually present, rarely absent | → 5 |
4. Androgynophore absent; nectaries on sepals | c. Tilioideae |
4. Androgynophore present or absent; nectaries on petals or androgynophore | d. Grewioideae |
5. Staminodes usually present; anthers 2- or 3-thecate | e. Byttnerioideae |
5. Staminodes absent or relatively small; anthers 1-thecate | f. Malvoideae |
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