Sidalcea covillei |
Sidalcea hirtipes |
|
---|---|---|
Owens Valley checkerbloom, Owens Valley sidalcea |
bluff mallow, bristly-stem checker-mallow, bristly-stem checkerbloom, hairy-stem checkermallow |
|
Habit | Herbs, perennial, 0.2–0.6 m, often glaucous, with fleshy, simple to clustered roots, without caudex or rhizomes. | Herbs, perennial, usually in colonies, 0.7–1.3(–1.8) m, not glaucous, with thick, rather woody taproot and coarse, elongate (cordlike) rhizomes 20–100 × 5 mm. |
Stems | several, clustered, erect, solid, often glaucous proximally, base sparsely, finely to coarsely stellate-hairy or hispid, hairs smaller distally. |
several, scattered, erect, arising from rhizome apices, usually slightly hollow, densely, harshly bristly-hirsute, hairs stiff, pustular, simple, forked, or stellate, often 2–2.5 mm. |
Leaves | 2–5 per stem, mostly basal; stipules linear-lanceolate, 3–6 × 1 mm; petiole (4–)5–10 cm, reduced on cauline leaves, proximal 1–4 times as long as blade, distal 1/2 times to as long as blade; blade fleshy, glaucous, rather densely stellate-hairy, proximalmost usually shallowly to deeply, ternately, palmately 5–7-lobed, 1.5–3 × 1.5–4 cm, lobes obovate, margins crenate-dentate, distal deeply 3–7-lobed, lobes linear, distalmost 2–4 cm wide. |
basal and cauline; stipules linear-subulate, 6–8 × 1–1.5 mm; petioles of proximal leaves 20–30 cm, 3 times as long as blades, reduced distally to 1/2 times as long as blades; blades: basal and proximalmost orbiculate to reniform, shallowly 5–9-lobed, 10–15 × 10–15 cm, thick, base narrowly cordate, lobe margins coarsely crenate-dentate, apex rounded, surfaces coarsely hirsute, hairs stiff; distal orbiculate, deeply palmately 5–7-lobed, incised ± to base, lobes sometimes lobed again, base cuneate, apex acute, deeply 2–3-toothed, surfaces long-hirsute or with 2–4-rayed, stellate hairs abaxially. |
Inflorescences | erect, open, calyces not conspicuously overlapping except sometimes in bud, branched or unbranched, nearly scapose, often 20+-flowered, slender, elongate, 1-sided or not, 6–30 cm; bracts inconspicuous, linear, 2-fid, 2–4 mm, shorter than calyx and pedicels. |
erect, spiciform to subcapitate, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, proximals usually long-pedunculate, unbranched or branched, 20+-flowered, 10+ flowers usually open on spike at same time, not interrupted, not 1-sided, usually to 8 cm, usually not elongate but sometimes slightly elongated in some populations and to 20 cm in fruit; bracts paired or single, linear, distal undivided, proximal distinct to base, 6 mm, mostly slightly longer than pedicels. |
Pedicels | 2–8(–10) mm; involucellar bractlets absent. |
1–3(–5) mm; involucellar bractlets absent. |
Flowers | bisexual; calyx 5–8 mm, uniformly, densely stellate-puberulent or few with longer rays; petals pale pink-lavender, veins paler, 10–15 mm; staminal column 4–5 mm, hairy; anthers white; stigmas 5 or 6. |
bisexual or unisexual and pistillate, plants gynodioecious; calyx often purple tinted, 9–11 mm, to 11–16 mm in fruit, margins ciliate, hairs 1–2 mm, surfaces finely stellate-hairy at base and with coarser, longer, simple and stellate hairs apically; petals usually pale pink to rose-lavender, rarely white, slightly or not pale-veined, (9–)10–21 mm, pistillate often 9–14 mm; staminal column 5–7(–10) mm, hairy; anthers white; stigmas 5–10. |
Seeds | 1 mm. |
2.5–3.5 mm. |
Schizocarps | 5 mm diam.; mericarps 5 or 6, 2.5 mm, sparsely glandular-puberulent, roughened, back reticulate-veined, sides strongly so, not pitted, mucro 0.1–0.3 mm. |
7–8 mm diam.; mericarps 5–10, 3.5–4 mm, glabrous or sparsely stellate-puberulent, roughened, prominently reticulate-veined, sides rugose and pitted, back less so, mucro 0.6–0.8 mm. |
2n | = 20. |
= 60. |
Sidalcea covillei |
Sidalcea hirtipes |
|
Phenology | Flowering (Apr–)May–Jun. | Flowering (Apr–)May–Jul(–Aug). |
Habitat | Alkaline flats, springs, meadows | Prairie remnants, coastal bluffs, open shrublands, fencerows, meadows, usually mesic, basaltic soil |
Elevation | 1100–1400 m (3600–4600 ft) | 0–200(–1200) m (0–700(–3900) ft) |
Distribution |
CA |
OR; WA
|
Discussion | Sidalcea covillei is one of two species of Sidalcea (along with S. neomexicana) with fleshy roots and adapted to alkaline conditions on flats. Its range (Owens Valley in Inyo County) and specialized habitat have made it vulnerable to any lowering of the water table and to grazing; it is listed as endangered in California. Most individuals of it were destroyed by construction of the Haiwee Reservoir. Once thought to have been extirpated, it was subsequently rediscovered. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Sidalcea hirtipes is uncommon and known from Clatsop, Lincoln, and Tillamook counties in Oregon and Clark, Lewis, and Wahkiakum counties in Washington. Its elevation and habitat vary, and it seems as much at home on steep coastal cliffs as in more inland, historic prairies and mountain meadows. Populations can appear to be large because of the long-rhizomatous and clonal nature of the plants; they are few and local. It is threatened by grazing, loss of habitat, fire suppression, road construction and maintenance, and changes in hydrology. It is a candidate for listing in Oregon and has been listed as endangered in Washington. Sidalcea hirtipes is characterized by its coarse indument of bristle hairs, its generally compact spikelike inflorescences, its relatively few, large, erect, hirsute leaves, and, especially, its extensive, coarse rhizomes. The inflorescences in some populations are elongated in fruit; its range, hirsute indument, and thick leaves along with coarse rhizomes help to distinguish it from other species. Stem internode length varies depending on habitat, as in many other Sidalcea. Molecular data suggest a relationship among S. hirtipes and S. asprella, S. celata, and S. gigantea (K. Andreasen and B. G. Baldwin 2003). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 329. | FNA vol. 6, p. 339. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. neomexicana var. covillei | |
Name authority | Greene: Cybele Columb. 1: 35. (1914) | C. L. Hitchcock: Perenn. Sp. Sidalcea, 42. (1957) |
Web links |