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dwarf checkerbloom, Redding checkerbloom

Oregon checker-mallow, Oregon checkerbloom

Habit Herbs, perennial, 0.4–0.8(–1) m, sometimes glaucous, with caudex and thick, woody rootstocks, not freely rooting, without rhizomes. Herbs, perennial, (0.3–)0.4–1.5 m, glaucous or not, with short, thick, rather woody taproot and branching caudex, without rhizomes or rhizomelike rootstocks (subsp. valida sometimes with rhizomes).
Stems

mostly single, usually erect, sometimes branched distally, solid (sometimes hollow in older, robust plants), sometimes glaucous, proximally densely bristly-hairy, hairs reflexed, stiff, simple, sometimes also coarse, 2–3-rayed, stellate, 1–1.6(–2.5) mm, pustulate, sparser distally, with minute, stellate hairs in inflorescence.

single or clustered, erect, rarely rooting at base, unbranched or distally branched, proximally usually solid, sometimes hollow in age, base glabrous, coarsely stellate-hairy to long-bristly, or glabrate, hairs usually becoming appressed, simple or stellate.

Leaves

mostly basal, some cauline;

stipules usually early-deciduous, linear-lanceolate, 3–7 × 1 mm;

proximal petioles 15–18 cm, 3–4 times as long as blade, distal usually to 1/2 times to as long as blade;

blades: basal rounded, usually palmately 7-lobed, shallowly incised, 4–8 × 4–8 cm, lobes 1.5–2 cm wide, margins crenate-dentate, 3–7-toothed, apices obtuse;

distal 3–10, palmately 5-lobed, deeply incised, 2–6 × 2–6 cm, lobes narrow, linear, 2–3.5 × 2–4 mm, margins entire or 1–5-toothed, surfaces: abaxial stellate-hairy, hairs 6-rayed, 1.5 mm, adaxial hairy, hairs simple, 1.5 mm, distalmost glabrate adaxially and sometimes simple-hairy.

basal and cauline, basal sometimes deciduous, cauline 3+;

stipules usually deciduous, linear to lanceolate, 4–6(–14) × 0.5–1(–2) mm;

petioles of basal and proximal leaves (5–)7–10(–35) cm, 3–5 times as long as blades, reduced distally, distalmost leaves sometimes subsessile;

blade cordate or reniform-orbiculate, 3–10(–15) × 3–10(–15) cm, base cordate, apex rounded, lobe apex often acute, surfaces glabrous or sparsely hairy, hairs minute, simple, forked, or stellate, proximal usually shallowly palmately 5–7(–9)-lobed, sometimes unlobed with margins crenate;

midstem more deeply (3–)5–9-lobed, lobes again palmately or pinnately lobed, distalmost unlobed or 3(–5)-lobed, segments unlobed or deeply lobed, narrow.

Inflorescences

erect, open, calyces not conspicuously overlapping except sometimes in bud, unbranched or branched, (5–)10–12(–23)-flowered, elongate, sometimes 1-sided, 10–40 cm;

bracts lanceolate, usually undivided, 3–4 mm, shorter than pedicels and calyx.

erect, usually spiciform, sometimes subcapitate, congested in bud, dense, calyces sometimes conspicuously overlapping in flower and sometimes in fruit, to open and elongate, few-branched or unbranched, 10–20+-flowered, flowers opening and closing sequentially from base to apex, sometimes 3–10 open on same day, not leafy-bracted, not 1-sided, (1.5–)10–30 cm, elongating in flower or fruit;

bracts linear to linear-lanceolate, undivided to 2-fid, proximal sometimes divided to base, 4–6(–7) mm, sometimes exceeding flower buds, usually equaling or longer than pedicels, shorter than calyx.

Pedicels

(3–)5–10(–15) mm;

involucellar bractlets absent.

1–3(–10) mm;

involucellar bractlets absent.

Flowers

usually bisexual, infrequently unisexual and pistillate, plants gynodioecious;

calyx (7–)9–10 mm, to 12–14 mm in fruit, densely stellate-puberulent;

petals pale pinkish lavender, often pale-veined especially when dry, pistillate 10–20 mm, bisexual 20–25(–31) mm;

staminal column 7–8(–10) mm, sparsely puberulent;

anthers white;

stigmas (6)7(8).

bisexual or unisexual and pistillate, plants gynodioecious;

calyx usually green, 3.5–10 mm, usually lightly reticulate-veined, glabrous or densely, uniformly stellate-puberulent or bristly, surface often obscured;

petals usually overlapping, pink or pink-lavender to dark rose-pink or magenta, not notably pale-veined, pistillate 5–10 mm, bisexual 8–15(–20) mm;

staminal column 4–6(–9) mm, hairy;

anthers white to pinkish;

stigmas 6–9.

Seeds

2.5 mm.

1.5–2.5 mm.

Schizocarps

6–8 mm diam.;

mericarps (6)7(8), 3–4 mm, minutely glandularpuberulent, glabrescent, margins sometimes sharpedged, roughened, strongly reticulate-veined, sides and back deeply pitted, honeycomblike, median line on back but not furrowed, mucro 1 mm.

4–7 mm diam.;

mericarps 6–9, 2–3 mm, sparsely glandular-puberulent, sometimes glabrous, not stellate-hairy, back and margins rounded, smooth or slightly reticulate-veined or pitted, infrequently prominently roughened at least on margins and/or back, mucro 0.1–0.7 mm.

2n

= 60.

= 20, 40, 60.

Sidalcea celata

Sidalcea oregana

Phenology Flowering May–Jun(–Aug).
Habitat Open oak woodlands, wet sites, sometimes on serpentine
Elevation 100–400(–1600) m (300–1300(–5200) ft)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; UT; WA; WY; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Sidalcea celata has been recognized as a local species endemic to Shasta and Tehama counties (especially near Redding) in the northern inner North Coast Ranges; it was previously included within S. malviflora. Hitchcock’s concept of S. celata as a subspecies was broader and included plants from additional counties that do not match the type very well; most have been re-identified as S. asprella or S. oregana. Sidalcea celata has been confused with S. asprella, S. gigantea, and S. robusta and appears to be variable. Its narrow distribution, preference for serpentine, typical lack of elongated rhizomes, basally retrorsely hirsute stems, presence of basal leaves, and relatively large bisexual flowers and fruits are distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Sidalcea oregana is variable and parts of it have been treated as distinct species, subspecies, varieties, or extremes of a continuum. The plants are generally characterized by their strictly erect, leafless inflorescences that are congested in bud, their variable, sparsely hairy, lobed leaves that are both basal and cauline, and their usual lack of rhizomes. They often have been characterized and distinguished from the S. malviflora group by smooth mericarps; this feature depends upon the subspecies and is not true of all plants of S. oregana. These are usually mountain plants; some grow at lower elevations toward the northern parts of the range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Primary peduncles (distal to distalmost leaves) longer than mature inflorescence; inflorescences dense, short-spiciform or subcapitate, 1.5–5 cm in age; calyces usually stellate-puberulent with sparse or no bristles; usually wetlands; California, Nevada
→ 2
1. Primary peduncles usually equaling or shorter than mature inflorescence; inflorescences open or dense, spiciform, 3–30 cm in age; calyces stellate-puberulent, with or without longer bristle hairs 0.5–2.5 mm; meadows, prairies, streamsides, not characteristically wetlands; British Columbia to California, Montana, Nevada, Utah, Wyoming
→ 3
2. Stems to 5 mm diam. near base, base stellate-bristly or hirsute, bristle hairs 2 mm; inflorescences 1.5–2.5 cm; 1100–2300 m.
subsp. hydrophila
2. Stems 5–10 mm diam. near base, base usually bristly-hirsute, sometimes with minute stellate hairs, bristle hairs 1–1.5 mm; inflorescences 2–5 cm; 100–200 m.
subsp. valida
3. Calyces to 8–13 mm in fruit, bristly-tomentose, some hairs 1.5–2.5 mm; Humboldt County, California.
subsp. eximia
3. Calyces to 3.5–7(–10) mm in fruit, short-stellate-hairy, sometimes also with bristles but not bristly-tomentose; British Columbia to California, Montana, Nevada, Utah, Wyoming
→ 4
4. Stem bases glabrous or stellate-hairy, hairs usually to 1 mm, sometimes bristly with appressed hairs, infrequently spreading; calyces usually uniformly stellate-puberulent; bracts usually shorter than young flower buds; rachis apex usually blunt or rounded.
subsp. oregana
4. Stem bases usually softly bristly-hirsute with hairs 1–2 mm or stellate-hairy, sometimes glabrous; calyces usually densely stellate-hairy, sometimes bristly; bracts usually longer than young flower buds; rachis apex usually acute.
subsp. spicata
Source FNA vol. 6, p. 329. FNA vol. 6, p. 347.
Parent taxa Malvaceae > subfam. Malvoideae > Sidalcea Malvaceae > subfam. Malvoideae > Sidalcea
Sibling taxa
S. asprella, S. calycosa, S. campestris, S. candida, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. oregana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
S. asprella, S. calycosa, S. campestris, S. candida, S. celata, S. covillei, S. cusickii, S. diploscypha, S. elegans, S. gigantea, S. glaucescens, S. hartwegii, S. hendersonii, S. hickmanii, S. hirsuta, S. hirtipes, S. keckii, S. malachroides, S. malviflora, S. multifida, S. nelsoniana, S. neomexicana, S. pedata, S. ranunculacea, S. reptans, S. robusta, S. setosa, S. sparsifolia, S. stipularis, S. virgata
Subordinate taxa
S. oregana subsp. eximia, S. oregana subsp. hydrophila, S. oregana subsp. oregana, S. oregana subsp. spicata, S. oregana subsp. valida
Synonyms S. malviflora var. celata, S. malviflora subsp. celata Sida oregana
Name authority (Jepson) S. R. Hill: Madroño 56: 106. (2009) (Nuttall ex Torrey & A. Gray) A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 20. (1849)
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