Sidalcea celata |
Sidalcea gigantea |
|
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dwarf checkerbloom, Redding checkerbloom |
giant checkerbloom |
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Habit | Herbs, perennial, 0.4–0.8(–1) m, sometimes glaucous, with caudex and thick, woody rootstocks, not freely rooting, without rhizomes. | Herbs, perennial, (0.8–)2(–2.5) m, not glaucous except on stems, usually in colonies, with rhizomes to 40–60 cm × (6–)10 mm, glabrescent with reflexed-appressed bristle hairs 2.5 mm. |
Stems | mostly single, usually erect, sometimes branched distally, solid (sometimes hollow in older, robust plants), sometimes glaucous, proximally densely bristly-hairy, hairs reflexed, stiff, simple, sometimes also coarse, 2–3-rayed, stellate, 1–1.6(–2.5) mm, pustulate, sparser distally, with minute, stellate hairs in inflorescence. |
single, scattered, erect, usually purple tinted proximally, hollow especially towards base, pithy distally, 10–14 mm diam. just distal to base, often glaucous, proximally densely retrorsely bristly-hairy and stellate-hairy, hairs pustulate, 1.5–2.5 mm, distally sometimes glabrous. |
Leaves | mostly basal, some cauline; stipules usually early-deciduous, linear-lanceolate, 3–7 × 1 mm; proximal petioles 15–18 cm, 3–4 times as long as blade, distal usually to 1/2 times to as long as blade; blades: basal rounded, usually palmately 7-lobed, shallowly incised, 4–8 × 4–8 cm, lobes 1.5–2 cm wide, margins crenate-dentate, 3–7-toothed, apices obtuse; distal 3–10, palmately 5-lobed, deeply incised, 2–6 × 2–6 cm, lobes narrow, linear, 2–3.5 × 2–4 mm, margins entire or 1–5-toothed, surfaces: abaxial stellate-hairy, hairs 6-rayed, 1.5 mm, adaxial hairy, hairs simple, 1.5 mm, distalmost glabrate adaxially and sometimes simple-hairy. |
mostly cauline, basal usually absent; stipules deciduous, with pink band on stem at base, narrowly lanceolate, (3.5–)5(–8) × 0.7(–1.5) mm; petioles of proximal leaves 6–8 cm, those of midstem leaves 10–14 cm, 2 times to as long as blade, apex curved with swollen portion or pulvinus 5–6 × 1.8 mm; blades of proximalmost cauline leaves rounded, shallowly, palmately 4- or 5-lobed, 0.5–0.7 × 0.5–0.7 cm, lobe margins irregularly, sparsely dentate, apices rounded, mid-cauline blades 5–7-lobed, 6.5–12 × 10–13 cm, wider than long, gradually reduced distally, lobes straplike, divided 3/4 to base, lobes 5.5 × 1.5–1.8 cm, margins coarsely dentate, surfaces sparsely, minutely hispid and stellate-puberulent, distal leaves deeply 5-lobed, otherwise similar in shape, leaves immediately below inflorescence greatly reduced, irregularly 2- or 3-lobed. |
Inflorescences | erect, open, calyces not conspicuously overlapping except sometimes in bud, unbranched or branched, (5–)10–12(–23)-flowered, elongate, sometimes 1-sided, 10–40 cm; bracts lanceolate, usually undivided, 3–4 mm, shorter than pedicels and calyx. |
erect, spiciform, open, calyces usually not overlapping, usually branched, branches 14–18 cm, each branch 10–20 flowered, not 1-sided, elongate, 5–20 cm; bracts 1, usually undivided, narrowly lanceolate, often 2-fid, 2.5 mm, equaling or slightly longer than pedicel, much shorter than calyx. |
Pedicels | (3–)5–10(–15) mm; involucellar bractlets absent. |
2–3(–5) mm; involucellar bractlets absent. |
Flowers | usually bisexual, infrequently unisexual and pistillate, plants gynodioecious; calyx (7–)9–10 mm, to 12–14 mm in fruit, densely stellate-puberulent; petals pale pinkish lavender, often pale-veined especially when dry, pistillate 10–20 mm, bisexual 20–25(–31) mm; staminal column 7–8(–10) mm, sparsely puberulent; anthers white; stigmas (6)7(8). |
bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–6 mm, to 8 mm in fruit, uniformly, densely stellate-puberulent, surface obscured; petals pale pink, pale-veined, pistillate 7–9 mm, bisexual (10–)14–20(–25) mm; staminal column 6–8 mm, stellate-puberulent; anthers white to cream or pale yellow; stigmas (6)7 or 8. |
Seeds | 2.5 mm. |
1.5 mm. |
Schizocarps | 6–8 mm diam.; mericarps (6)7(8), 3–4 mm, minutely glandularpuberulent, glabrescent, margins sometimes sharpedged, roughened, strongly reticulate-veined, sides and back deeply pitted, honeycomblike, median line on back but not furrowed, mucro 1 mm. |
6–7 mm diam.; mericarps (6)7 or 8, sides 3 mm, thick, back and margins thick, rounded, reticulate-veined, pitted, back with prominent groove, top sparsely glandular-stellate-puberulent, mucro 1 mm. |
2n | = 60. |
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Sidalcea celata |
Sidalcea gigantea |
|
Phenology | Flowering May–Jun(–Aug). | Flowering Jul–Sep. |
Habitat | Open oak woodlands, wet sites, sometimes on serpentine | Moist to wet, forested slopes, seeps, stream margins, meadows, coniferous forests |
Elevation | 100–400(–1600) m (300–1300(–5200) ft) | (600–)900–1700 m ((2000–)3000–5600 ft) |
Distribution |
CA |
CA |
Discussion | Sidalcea celata has been recognized as a local species endemic to Shasta and Tehama counties (especially near Redding) in the northern inner North Coast Ranges; it was previously included within S. malviflora. Hitchcock’s concept of S. celata as a subspecies was broader and included plants from additional counties that do not match the type very well; most have been re-identified as S. asprella or S. oregana. Sidalcea celata has been confused with S. asprella, S. gigantea, and S. robusta and appears to be variable. Its narrow distribution, preference for serpentine, typical lack of elongated rhizomes, basally retrorsely hirsute stems, presence of basal leaves, and relatively large bisexual flowers and fruits are distinctive. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea gigantea is likely the tallest Sidalcea species; it can be distinguished also by its range (high Cascades and the northern high Sierra Nevada), retrorse hirsute stem bases, thick, hollow stems, and massive, hirsute rhizome systems. Most large patches appear to be clonal and generally produce either bisexual or unisexual, pistillate stems. Sidalcea gigantea is closely related to, and has been confused with, both S. asprella and S. celata. Its leaves are most like those of S. asprella in that they are usually similar in shape throughout the stem; the tall stems and massive rhizome systems distinguish it from S. asprella, as do its occurrence at higher elevations and its later flowering time. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 329. | FNA vol. 6, p. 332. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. malviflora var. celata, S. malviflora subsp. celata | |
Name authority | (Jepson) S. R. Hill: Madroño 56: 106. (2009) | G. L. Clifton: Madroño 56: 285, figs. 1 – 3. (2010) |
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