Sidalcea campestris |
Sidalcea cusickii |
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meadow checker-mallow, meadow checkerbloom, meadow sidalcea |
Cusick's checkerbloom, Cusick's checkermallow, Cusick's sidalcea |
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Habit | Herbs, perennial, 0.5–2 m, sometimes glaucous in age, with thick, rather woody taproot and short rhizomes usually to 10 cm × 10 mm. | Herbs, perennial, 0.4–1.8 m, not glaucous, with thick taproot and freely-rooting rhizomes usually 3–10 mm diam. |
Stems | single or clustered, erect, base often decumbent-ascending, solid (proximally hollow on older stems), proximally densely bristly-hirsute, hairs simple or forked, 0.5–1 mm, sometimes mixed with minute, stellate hairs, sometimes glabrous and glaucous in age, hairs usually more appressed, simple, stellate, or sparse distally. |
usually several, clustered, erect, often purplish, often thick and hollow proximally, usually proximally ± scabrous, hairs stellate and distally stellate-hairy, sometimes glabrous. |
Leaves | mostly cauline; stipules lanceolate, 4–11 × 1–1.5 mm; petioles of proximal leaves 15–20 cm, 2–3 times as long as blades, distal reduced to 0.5–10 cm, 1/2 times to as long as blades; blade 10–15 × 10–15 cm, smaller distally, surfaces scabrid-hairy, hairs dense, simple or stellate, stiff, stellate hairs mostly on adaxial surfaces, proximal blades orbiculate, shallowly to deeply palmately 7–9-lobed, 5–15 × 5–15 cm, base cordate, margins coarsely crenate-serrate, lobes apically 2–5-toothed; distal cauline leaves variable, mid blades usually palmately divided nearly to base into 5–7 cuneate lobes, 15 × 15 cm, lobes deeply cut to laciniately dissected, distal blades divided into (3–)5–7 linear, marginally subentire segments, surfaces glabrescent or with few hairs on abaxial surface veins, ciliate. |
mostly cauline; stipules persistent or deciduous, lanceolate, 6–8 × 1–1.5 mm; proximal petioles 25–30 cm, 4–5 times as long as blade, reduced to 7–13 cm on midstem leaves and to 2–3 cm on distalmost leaves, these to 1/2 times as long as blades; basal blades wide-ovate to orbiculate, shallowly to deeply, palmately 5–7(–9)-lobed, 6–13 × 6–13 cm, lobe margins toothed, teeth rounded, sometimes Ribes-like, base cordate, apex rounded, surfaces: abaxial densely stellate-pubescent, adaxial glabrous or sparsely stellate-pubescent on veins; cauline blades round, divided nearly to base, 5–9-lobed, lobes again deeply incised to subentire, 10–20 × 10–20 cm, distalmost 5–7-cleft to base. |
Inflorescences | erect, open, spiciform, calyces not conspicuously overlapping except sometimes in bud, often branched from distal leaves, 15+-flowered, elongate, not 1-sided, 10–35 cm, proximal flowers spaced several cm apart, not leafy-bracted; bracts linear, distal undivided or 2-fid, proximal separate to base, 4–10 mm, usually equaling or longer than pedicels. |
erect, spiciform, dense, calyces usually conspicuously overlapping in flower and sometimes in fruit, usually 20–30-branched per primary stem, 20+-flowered, elongate, not 1-sided, most flowers usually open at same time, branches relatively short, each spike unit 6 cm, longer in fruit; bracts ovate-lanceolate to subulate, undivided, 1–6(–10) mm, subequal or longer than pedicels, much shorter than calyx. |
Pedicels | 3–6(–20) mm; involucellar bractlets absent. |
1–2(–5) mm; involucellar bractlets absent. |
Flowers | bisexual or unisexual and pistillate, plants gynodioecious; calyx 5–9 mm, pistillate 5–7 mm, bisexual 6–9 mm, 8–10 mm in fruit, uniformly, densely stellate-hairy or with coarser, longer, stellate hairs to 2 mm; petals usually not overlapping, nearly white to pale pink or pale lavender, pistillate 9–12 mm, bisexual 13–25 mm; staminal column 5–7 mm, hairy; anthers white to pale pink; stigmas (6 or)7 or 8. |
bisexual or unisexual and pistillate, pistillate plants more frequent than bisexual ones, some flowers may be staminate, plants gynodioecious; calyx urceolate with swollen base especially in young fruit, pistillate 6–8 mm, bisexual 6–10 mm, strongly reticulate-veined, usually finely stellate-puberulent, sometimes glabrate, lobes usually purple tinted; petals 7 or 8, pale pink, pinkish rose, or rose-purple, usually neither pale-veined nor white at base, pistillate 8–12(–14) mm, staminate or bisexual (10–)11–19(–23) mm; staminal column 5–6 mm, hairy; anthers white; stigmas 7 or 8. |
Seeds | 2.5 mm. |
2 mm. |
Schizocarps | 7–8 mm diam.; mericarps (6 or)7 or 8, 3.5 mm, roughened, sides prominently reticulate-rugose and pitted, back less so and glandular-puberulent, mucro 0.5–1 mm. |
6 mm diam.; mericarps 7 or 8, 3 mm, apical margins ± rounded, sides smooth or slightly reticulate-veined, not pitted, back essentially smooth, very sparsely glandular-puberulent near tip, mucro 0.5–1 mm. |
2n | = 60. |
= 20. |
Sidalcea campestris |
Sidalcea cusickii |
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Phenology | Flowering May–Aug. | Flowering Jun–Aug. |
Habitat | Open shrublands, meadows, hedgerows, prairies | Moist to wet, mostly black, adobe soil, lowland and mountain meadows, often with Juncus and Camassia |
Elevation | 40–200 m (100–700 ft) | 100–200(–500), 1000–1400 m (300–700(–1600), 3300–4600 ft) |
Distribution |
OR
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OR
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Discussion | Sidalcea campestris is one of the taller species of Sidalcea and can be distinguished also by its range, hirsute indument, long inflorescences with spaced, long-stalked flowers, narrow very pale petals, and deeply incised pinnatifid stem leaves. It has been confused with S. hirtipes, S. nelsoniana, and S. oregana; it differs especially in characters of rhizomes, inflorescences, and leaves. It is a candidate for listing as threatened or endangered in Oregon. Specimens from British Columbia and Washington identified as S. campestris are either S. campestris escaped from cultivation or S. hendersonii. It is known as a native only from the Willamette Valley area (Multnomah and Washington to Benton and Linn counties). Apparently, it was introduced near Seattle, Washington; it may not persist there. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Sidalcea cusickii is showy and distinctive. It has been confused with S. campestris, S. hendersonii, S. nelsoniana, S. oregana (subsp. spicata), S. setosa, and S. virgata. Molecular data support a close relationship with S. hendersonii and S. virgata (K. Andreasen and B. G. Baldwin 2003b). Hitchcock distinguished subsp. cusickii and subsp. purpurea, the latter with purple, ciliate calyx lobes and stem proximally glabrous and thought to be more closely related to S. hendersonii and S. nelsoniana. The two variations overlap in range and characteristics and are not recognized here. There appear to be two groups of populations at different elevation ranges with little overlap. Overall, S. cusickii can be distinguished by its range and by its spikelike, many-flowered, thick, compounded racemes in which essentially all of the flowers are open at the same time. It is also unusual in the preponderance of pistillate-flowered individuals in a given population and the presence of what appear to be truly staminate, rather than bisexual, flowers on some individuals. It is found in the Willamette and Umpqua valley region in Douglas, Josephine, and Lane counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 328. | FNA vol. 6, p. 330. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | S. asplenifolia, S. sylvestris | S. cusickii subsp. purpurea, S. oregana var. cusickii |
Name authority | Greene: Bull. Calif. Acad. Sci. 1: 76. (1885) | Piper: Proc. Biol. Soc. Wash. 29: 99. (1916) |
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