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strong bladderpod

Snake River or creeping twinpod, Snake River twinpod

Habit Perennials; caudex branched, (thickened); densely pubescent, trichomes (sessile), several-rayed, rays furcate or bifurcate, usually fused toward base, (strongly tuberculate). Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout).
Stems

several from base, erect or outer decumbent, (unbranched), to 2 dm.

several from base, ± erect, exceeding basal rosette by ± 0.5 dm.

Basal leaves

blade elliptic to lanceolate or obovate, 3–8 cm, margins entire.

(forming a strong rosette; long-petiolate);

blade oblanceolate to ovate or orbicular, (1.5–)2–4(–8) cm, (base usually abruptly tapering to petiole), margins entire.

Cauline leaves

(proximal shortly petiolate or sessile, distal sessile);

blade elliptic or obovate, to 2 cm, margins entire.

blade oblanceolate, 1–2 cm, margin entire, (apex acute).

Racemes

dense.

congested, (greatly exceeding leaves).

Flowers

sepals narrowly elliptic or oblong, 4.5–5.3 mm, (tapering to the somewhat thickened, cucullate apex, lateral pair subsaccate);

petals (bright yellow), lingulate or broadly obovate, 7.5–8.5 mm, (narrowing to broad claw, joining in an arch, margins lacerate).

sepals often keeled, 6–8 mm;

petals spatulate, 8–10 mm.

Fruiting pedicels

(divaricate-ascending to horizontal, straight to loosely curved), to 15 mm.

(spreading, straight or slightly curved), 7–11 mm.

Fruits

(sessile or substipitate), suborbicular to broadly ovate or ellipsoid, slightly compressed, 6–8 mm;

valves pubescent;

ovules 12–22 per ovary;

style 2–3 mm.

didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed;

replum linear to oblong, as wide as or wider than fruit;

ovules 8 per ovary;

style 7–9 mm.

Seeds

flattened.

flattened.

2n

= 10.

= 16.

Physaria valida

Physaria integrifolia

Phenology Flowering Apr–May. Flowering (May-)Jun–Jul(-Aug).
Habitat Limestone soils, steep slopes, roadcuts, open woods Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale
Elevation 1900-2200 m (6200-7200 ft) 1900-2700 m (6200-8900 ft)
Distribution
from FNA
NM; TX
[BONAP county map]
from FNA
ID; MT; WY
[BONAP county map]
Discussion

Physaria valida is known from the Sacramento and White mountains of south central New Mexico, and southward through the Guadalupe Mountains to Hudspeth County, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 664. FNA vol. 7, p. 644.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella valida, Lesquerella lepidota P. didymocarpa var. integrifolia, P. integrifolia var. monticola
Name authority (Greene) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) (Rollins) Lichvar: Madroño 31: 203. (1984)
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