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strong bladderpod

Engelmann's bladderpod

Habit Perennials; caudex branched, (thickened); densely pubescent, trichomes (sessile), several-rayed, rays furcate or bifurcate, usually fused toward base, (strongly tuberculate). Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate).
Stems

several from base, erect or outer decumbent, (unbranched), to 2 dm.

few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm.

Basal leaves

blade elliptic to lanceolate or obovate, 3–8 cm, margins entire.

blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent).

Cauline leaves

(proximal shortly petiolate or sessile, distal sessile);

blade elliptic or obovate, to 2 cm, margins entire.

(proximal often petiolate, distal sessile or subsessile);

blade oblanceolate to linear, 1–4 cm, margins entire.

Racemes

dense.

dense, (subumbellate).

Flowers

sepals narrowly elliptic or oblong, 4.5–5.3 mm, (tapering to the somewhat thickened, cucullate apex, lateral pair subsaccate);

petals (bright yellow), lingulate or broadly obovate, 7.5–8.5 mm, (narrowing to broad claw, joining in an arch, margins lacerate).

sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate);

petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse).

Fruiting pedicels

(divaricate-ascending to horizontal, straight to loosely curved), to 15 mm.

(ascending), relatively short.

Fruits

(sessile or substipitate), suborbicular to broadly ovate or ellipsoid, slightly compressed, 6–8 mm;

valves pubescent;

ovules 12–22 per ovary;

style 2–3 mm.

(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20 per ovary;

style 3.5–5 mm.

Seeds

flattened.

flattened.

2n

= 10.

= 12, 24, 36.

Physaria valida

Physaria engelmannii

Phenology Flowering Apr–May. Flowering Apr–May.
Habitat Limestone soils, steep slopes, roadcuts, open woods Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops
Elevation 1900-2200 m (6200-7200 ft) 150-400 m (500-1300 ft)
Distribution
from FNA
NM; TX
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

Physaria valida is known from the Sacramento and White mountains of south central New Mexico, and southward through the Guadalupe Mountains to Hudspeth County, Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 664. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella valida, Lesquerella lepidota Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella
Name authority (Greene) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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