Physaria valida |
Physaria didymocarpa |
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strong bladderpod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; caudex branched, (thickened); densely pubescent, trichomes (sessile), several-rayed, rays furcate or bifurcate, usually fused toward base, (strongly tuberculate). | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||
Stems | several from base, erect or outer decumbent, (unbranched), to 2 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | blade elliptic to lanceolate or obovate, 3–8 cm, margins entire. |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | (proximal shortly petiolate or sessile, distal sessile); blade elliptic or obovate, to 2 cm, margins entire. |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | dense. |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals narrowly elliptic or oblong, 4.5–5.3 mm, (tapering to the somewhat thickened, cucullate apex, lateral pair subsaccate); petals (bright yellow), lingulate or broadly obovate, 7.5–8.5 mm, (narrowing to broad claw, joining in an arch, margins lacerate). |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (divaricate-ascending to horizontal, straight to loosely curved), to 15 mm. |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | (sessile or substipitate), suborbicular to broadly ovate or ellipsoid, slightly compressed, 6–8 mm; valves pubescent; ovules 12–22 per ovary; style 2–3 mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 10. |
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Physaria valida |
Physaria didymocarpa |
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Phenology | Flowering Apr–May. | |||||||||
Habitat | Limestone soils, steep slopes, roadcuts, open woods | |||||||||
Elevation | 1900-2200 m (6200-7200 ft) | |||||||||
Distribution |
NM; TX |
ID; MT; WA; WY; AB; BC
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Discussion | Physaria valida is known from the Sacramento and White mountains of south central New Mexico, and southward through the Guadalupe Mountains to Hudspeth County, Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 664. | FNA vol. 7, p. 633. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella valida, Lesquerella lepidota | Vesicaria didymocarpa, Coulterina didymocarpa | ||||||||
Name authority | (Greene) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||
Web links |