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Snake Range bladderpod

Columbia bladder-pod, Douglas' bladder-pod

Habit Perennials; caudex simple; densely pubescent, trichomes 5–7-rayed, rays bifurcate or trifurcate. Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout).
Stems

several from base, erect or outer usually decumbent toward base, (from below a terminal tuft of mostly erect leaves, unbranched), 1–2 dm.

simple from base, erect, (usually unbranched), to 4.5 dm.

Basal leaves

blade ovate to elliptic, 2–4 cm, (base gradually narrowed to petiole), margins entire, (surfaces densely pubescent with trichome layers).

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

Cauline leaves

(remote, proximal shortly petiolate);

blade spatulate to oblanceolate, similar to basal, (base often cuneate), margins entire.

similar to basal, blade narrowly linear or, sometimes, orbicular.

Racemes

elongated.

loose (lax).

Flowers

sepals linear-oblong, 5–7 mm, (lateral pair subsaccate);

petals (erect), lingulate, 8–10 mm, (claw barely differentiated from blade).

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

Fruiting pedicels

(recurved), 7–10 mm.

(recurved, straight, curved, or sigmoid), 6–20 mm.

Fruits

(usually pendent, sessile), subglobose, slightly flattened (angustiseptate), 4–5 mm;

valves densely pubescent, trichomes somewhat spreading;

ovules 8–12 per ovary;

style 4–5 mm.

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

Seeds

plump, (mucilaginous when wetted).

flattened.

2n

= 10, 30.

Physaria pendula

Physaria douglasii

Phenology Flowering May–Jun.
Habitat Limestone gravel and cobbles, typically with junipers
Elevation 1700-2100 m (5600-6900 ft)
Distribution
from FNA
NV
[BONAP county map]
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 657. FNA vol. 7, p. 634.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella pendula Lesquerella douglasii
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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