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Oregon twin-pod

frosty bladderpod, Pagosa bladderpod

Habit Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). Perennials; caudex simple or branched, (covered with persistent leaf bases); densely pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout).
Stems

several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm.

simple or several from base, decumbent or erect, (unbranched), to 2 dm.

Basal leaves

(petiole slender, usually incised or with broad teeth along petiole);

blade obovate, 4–6 cm, margins entire.

(petiole sharply differentiated from blade, slender);

blade suborbicular or obovate to rhombic, 4–8 cm, (base abruptly narrowed to petiole), margins entire, sinuate, or shallowly dentate, (abaxial surface densely pubescent, adaxial lightly pubescent).

Cauline leaves

blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute).

(proximal petiolate, distal sessile);

blade obovate to rhombic, 0.8–2.3 cm, margins entire or shallowly toothed.

Racemes

somewhat loose, (5–15 cm).

dense, (somewhat elongated in fruit).

Flowers

sepals oblong, 5–7 mm;

petals (lemon yellow), spatulate, 9–12 mm.

sepals elliptic or oblong, ca. 6 mm, (lateral pair not saccate or subsaccate, cucullate, median pair thickened apically, cucullate);

petals spatulate, ca. 9 mm, (claw expanded at base).

Fruiting pedicels

(spreading or ascending, curved, fruits not pendent), 10–20 mm.

(horizontal to ascending, sigmoid or slightly curved), 8–11 mm, (stout).

Fruits

didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading;

replum broadly lanceolate, as wide as or wider than fruit, apex acute;

ovules 8 per ovary;

style 1–2 mm.

(sessile or substipitate, often becoming copper-red in age), subglobose or ellipsoid, inflated, 6–9 mm, (firm, glossy);

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8(–12) per ovary;

style 3.5–7 mm.

Seeds

flattened.

somewhat flattened.

2n

= 8.

Physaria oregona

Physaria pruinosa

Phenology Flowering Apr–Jun. Flowering May–Jun(-Aug).
Habitat Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils Mancos slate or shale, meadows, gentle slopes, edges of ponderosa pine stands
Elevation 900-1900 m (3000-6200 ft) 2100-2600 m (6900-8500 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM
[BONAP county map]
Discussion

Of conservation concern.

The one New Mexico population is near the border with Colorado, in Rio Arriba County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 655. FNA vol. 7, p. 658.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Coulterina oregona, Lesquerella oregona Lesquerella pruinosa
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) (Greene) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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