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Oregon twin-pod

Newberry twinpod, Newberry's twinpod

Habit Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length.
Stems

several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm.

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

Basal leaves

(petiole slender, usually incised or with broad teeth along petiole);

blade obovate, 4–6 cm, margins entire.

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

Cauline leaves

blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute).

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

Racemes

somewhat loose, (5–15 cm).

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

Flowers

sepals oblong, 5–7 mm;

petals (lemon yellow), spatulate, 9–12 mm.

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

Fruiting pedicels

(spreading or ascending, curved, fruits not pendent), 10–20 mm.

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

Fruits

didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading;

replum broadly lanceolate, as wide as or wider than fruit, apex acute;

ovules 8 per ovary;

style 1–2 mm.

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

Seeds

flattened.

slightly flattened, (ovate).

2n

= 8.

Physaria oregona

Physaria newberryi

Phenology Flowering Apr–Jun.
Habitat Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils
Elevation 900-1900 m (3000-6200 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 655. FNA vol. 7, p. 652.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms Coulterina oregona, Lesquerella oregona P. didymocarpa var. newberryi
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861)
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