Physaria lindheimeri |
Physaria navajoensis |
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Lindheimer's bladderpod |
Navajo bladderpod |
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Habit | Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). |
Stems | several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm. |
several from base (crowded), erect, not exceeding leaves. |
Basal leaves | blade pinnatisect to repand, 3–9(–14) cm, margins entire. |
usually absent. |
Cauline leaves | (sometimes secund, proximal usually petiolate, distal sessile); blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate. |
(petiole not differentiated from blade); blade linear-oblanceolate, 3–8(–13) mm, margins entire. |
Racemes | dense. |
(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). |
Flowers | sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate); petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw). |
sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle). |
Fruiting pedicels | (horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm. |
(ascending to divaricate-ascending, straight), 3.5–6 mm. |
Fruits | globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth); valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules (8–)12–16(–20) per ovary; style (1.5–)2–3(–4) mm. |
(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4–8 per ovary; style 1.8–3 mm. |
Seeds | flattened. |
plump or slightly flattened, (strongly mucilaginous). |
2n | = 12. |
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Physaria lindheimeri |
Physaria navajoensis |
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Phenology | Flowering Dec–Apr. | Flowering May-early Jun. |
Habitat | Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies | Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone |
Elevation | 20-800 m (100-2600 ft) | 2200-2400 m (7200-7900 ft) |
Distribution |
TX; Mexico (Tamaulipas) |
AZ; NM |
Discussion | Of conservation concern. Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 651. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri | Lesquerella navajoensis |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |