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Lindheimer's bladderpod

Navajo bladderpod

Habit Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo).
Stems

several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm.

several from base (crowded), erect, not exceeding leaves.

Basal leaves

blade pinnatisect to repand, 3–9(–14) cm, margins entire.

usually absent.

Cauline leaves

(sometimes secund, proximal usually petiolate, distal sessile);

blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate.

(petiole not differentiated from blade);

blade linear-oblanceolate, 3–8(–13) mm, margins entire.

Racemes

dense.

(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves).

Flowers

sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate);

petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw).

sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate);

petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle).

Fruiting pedicels

(horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm.

(ascending to divaricate-ascending, straight), 3.5–6 mm.

Fruits

globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules (8–)12–16(–20) per ovary;

style (1.5–)2–3(–4) mm.

(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute);

valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not);

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

flattened.

plump or slightly flattened, (strongly mucilaginous).

2n

= 12.

Physaria lindheimeri

Physaria navajoensis

Phenology Flowering Dec–Apr. Flowering May-early Jun.
Habitat Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation 20-800 m (100-2600 ft) 2200-2400 m (7200-7900 ft)
Distribution
from FNA
TX; Mexico (Tamaulipas)
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 649. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri Lesquerella navajoensis
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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