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Lindheimer's bladderpod

intermountain bladderpod, skyline bladderpod

Habit Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). Perennials; caudex simple or branched (tightly); sparsely to densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays furcate or bifurcate, distinct or slightly fused at base, (sometimes umbonate, tuberculate throughout).
Stems

several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm.

few to several from base, decumbent, 0.5–1(–2) dm, (rather stout, sparsely pubescent).

Basal leaves

blade pinnatisect to repand, 3–9(–14) cm, margins entire.

blade elliptic to suborbicular, 1.5–3.5(–5.5) cm, margins entire or shallowly dentate (at base, surfaces densely pubescent, silvery).

Cauline leaves

(sometimes secund, proximal usually petiolate, distal sessile);

blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate.

(petiolate or distal nearly sessile);

blade elliptic to obovate, 0.5–1.5 cm, margins entire.

Racemes

dense.

dense, congested, (few-flowered).

Flowers

sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate);

petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw).

sepals lanceolate, oblanceolate, or narrowly elliptic, 3.8–5 mm, (median pair thickened apically, cucullate);

petals narrowly lanceolate to linear, 6–10(–13) mm.

Fruiting pedicels

(horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm.

(spreading or recurved, sometimes loosely sigmoid), 2–6.5 mm.

Fruits

globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules (8–)12–16(–20) per ovary;

style (1.5–)2–3(–4) mm.

(sessile or substipitate), broadly obcordate, obdeltate, or obcompressed, slightly compressed (angustiseptate), 3–5(–7) mm;

valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, trichomes appressed;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style (1.8–)3–6(–7) mm.

Seeds

flattened.

slightly flattened, (ellipsoid to suborbicular).

2n

= 12.

Physaria lindheimeri

Physaria hemiphysaria

Phenology Flowering Dec–Apr.
Habitat Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies
Elevation 20-800 m (100-2600 ft)
Distribution
from FNA
TX; Mexico (Tamaulipas)
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles 3-5 mm; fruiting pedicels 4-6.5 mm; fruit valves pubescent, often sparsely so.
subsp. hemiphysaria
1. Styles 1.8-2.5(-3) mm; fruiting pedicels 2-4(-5.5) mm; fruit valves glabrous or sparsely pubescent.
subsp. lucens
Source FNA vol. 7, p. 649. FNA vol. 7, p. 642.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hemiphysaria subsp. hemiphysaria, P. hemiphysaria subsp. lucens
Synonyms Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri Lesquerella hemiphysaria
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) (Maguire) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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