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Lindheimer's bladderpod

Devils Gate twinpod

Habit Annuals or biennials; with a fine taproot; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate at base, sometimes bifurcate, (rough-tuberculate throughout). Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth).
Stems

several from base, erect or outer decumbent, (often several-branched, branches slender and flexuous), to 8 dm.

simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm.

Basal leaves

blade pinnatisect to repand, 3–9(–14) cm, margins entire.

blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed).

Cauline leaves

(sometimes secund, proximal usually petiolate, distal sessile);

blade elliptic, 1–6 cm, (distal with cuneate base), margins entire or deeply dentate.

(2–4);

blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute).

Racemes

dense.

condensed.

Flowers

sepals elliptic to oblong, 3–5.5 mm, (median pair slightly thickened apically, cucullate);

petals (sometimes drying slightly purplish), suborbicular or broadly ovate, 4.5–7(–9) mm, (narrowing gradually to short claw).

sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median);

petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade).

Fruiting pedicels

(horizontal or recurved and ascending at tip, sometimes loosely sigmoid), (5–)10–20 mm.

(divaricate-ascending, nearly straight), 6–10 mm.

Fruits

globose or broadly ellipsoid, not or slightly inflated, (4–)5–8 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules (8–)12–16(–20) per ovary;

style (1.5–)2–3(–4) mm.

strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery);

valves (retaining seeds after dehiscence), pubescent;

replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse;

ovules 4–8 per ovary;

style 4–5 mm, (sparsely pubescent or glabrous).

Seeds

flattened.

plump.

2n

= 12.

Physaria lindheimeri

Physaria eburniflora

Phenology Flowering Dec–Apr. Flowering May–Jun.
Habitat Heavy, black, claylike soils, or lighter, sandy soils, thickets, field-margins, roadsides, coastal prairies Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus
Elevation 20-800 m (100-2600 ft) 1800-3000 m (5900-9800 ft)
Distribution
from FNA
TX; Mexico (Tamaulipas)
[BONAP county map]
from FNA
WY
[BONAP county map]
Source FNA vol. 7, p. 649. FNA vol. 7, p. 635.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria lindheimeri, Alyssum lindheimeri, Lesquerella gracilis var. pilosa, Lesquerella lindheimeri
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) Rollins: Brittonia 33: 333. (1981)
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