Physaria klausii |
Physaria chambersii |
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Rogers Pass or klaus' or divide bladderpod |
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (loosely spreading), 3–5-rayed, rays distinct, furcate (with exceptionally long branches). | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). |
Stems | simple from base, erect to decumbent, (slender), 0.6–1.5 dm. |
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
Basal leaves | blades obovate to deltate, 1.5–3(–4) cm, margins entire or outer one with 1 or 2 broad teeth. |
(petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
Cauline leaves | blade oblanceolate to spatulate, 0.6–1.5 cm, margins entire. |
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
Racemes | loose. |
congested. |
Flowers | sepals (green-yellow, often tinged with purple), elliptic, 3–4.6 mm; petals oblanceolate, 6–8 mm (claw expanded). |
sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (sigmoid), 5–9 mm. |
(divaricate, slightly sigmoid), 8–15 mm. |
Fruits | (depressed), broadly obovate, compressed (angustiseptate), 2–4 mm, (apex slightly bilobed to nearly truncate); valves densely pubescent, trichomes strongly ascending, spreading, long, (appearing fuzzy), pubescent inside; ovules 4 per ovary; style 3–4 mm, (pubescent or glabrous). |
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
Seeds | flattened. |
flattened. |
2n | = 8, 10, 16, 24. |
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Physaria klausii |
Physaria chambersii |
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Phenology | Flowering Jul. | Flowering Apr–Jul. |
Habitat | Open gravel slides, solifluction cross-stripes of shale rubble, barren shale-derived soil | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas |
Elevation | 1200-1900 m (3900-6200 ft) | 1500-3200 m (4900-10500 ft) |
Distribution |
MT |
AZ; CA; ID; NV; OR; UT
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Discussion | Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 647. | FNA vol. 7, p. 631. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella klausii | P. chambersii var. canaani, P. chambersii var. sobolifera |
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) |
Web links |