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Graham's twinpod

roundtip twinpod

Habit Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled).
Stems

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm.

Basal leaves

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse).

Cauline leaves

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute).

Racemes

loose, (elongated).

congested, (elongated in fruit).

Flowers

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

sepals oblong, 6–8 mm;

petals spatulate, to 10 mm.

Fruiting pedicels

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

(usually curving upward, sigmoid), 6–10 mm.

Fruits

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose;

replum oblong, often constricted, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

Seeds

plump, (suborbicular).

flattened.

2n

= 8, 16.

Physaria grahamii

Physaria vitulifera

Phenology Flowering May–Jun. Flowering Apr–Jun.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale
Elevation 2100-2900 m (6900-9500 ft) 1600-3000 m (5200-9800 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 642. FNA vol. 7, p. 665.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina
Synonyms P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda
Name authority C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) Rydberg: Bull. Torrey Bot. Club 28: 278. (1901)
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