Physaria grahamii |
Physaria tenella |
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Graham's twinpod |
little bladderpod, Moapa bladderpod |
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Habit | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. | Annuals or, rarely, biennials; with a taproot; densely pubescent, trichomes (simple or stellate, sessile or short-stalked), 4–7-rayed, rays usually furcate, rarely bifurcate, (nearly smooth to finely tuberculate). |
Stems | several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
several from base, decumbent to erect, (several-branched, frequently stout), 1.5–6 dm. |
Basal leaves | (outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
blade elliptic, (1.5–)3–6.5 cm, margins entire, repand, or shallowly dentate. |
Cauline leaves | similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
(proximal shortly petiolate, distal sessile); blade linear to elliptic or obovate, (0.5–)1–3.5(–4.5) cm, margins entire or repand. |
Racemes | loose, (elongated). |
loose. |
Flowers | sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
sepals oblong, lanceolate, or elliptic, (3–)3.5–6(–7.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (yellow to orange), suborbicular or obovate, (5–)6.5–8(–11) mm, (narrowing gradually to broad claw, usually widened at base). |
Fruiting pedicels | (ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
(recurved, sigmoid), 5–15 mm. |
Fruits | didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
(sessile or shortly stipitate), orbicular or obovoid, often slightly compressed, (3.5–)4–6 mm; valves sparsely pubescent, trichomes sessile and stellate, densely pubescent inside, trichomes simple or branched; ovules 4–12 per ovary; style 2–4.5 mm. |
Seeds | plump, (suborbicular). |
flattened. |
2n | = 10, 20. |
|
Physaria grahamii |
Physaria tenella |
|
Phenology | Flowering May–Jun. | Flowering Feb–May. |
Habitat | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay | Sandy soils, gravel, clayey loam, loose rocky slopes, washes, desert slopes and plains, lava hills, frequently in or near bushes |
Elevation | 2100-2900 m (6900-9500 ft) | (0-)600-1900 m ((0-)2000-6200 ft) |
Distribution |
UT |
AZ; CA; NV; UT; Mexico (Sonora)
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Discussion | Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 642. | FNA vol. 7, p. 663. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda | Lesquerella tenella, Lesquerella gordonii var. sessilis |
Name authority | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) |
Web links |