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Graham's twinpod

little bladderpod, Moapa bladderpod

Habit Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. Annuals or, rarely, biennials; with a taproot; densely pubescent, trichomes (simple or stellate, sessile or short-stalked), 4–7-rayed, rays usually furcate, rarely bifurcate, (nearly smooth to finely tuberculate).
Stems

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

several from base, decumbent to erect, (several-branched, frequently stout), 1.5–6 dm.

Basal leaves

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

blade elliptic, (1.5–)3–6.5 cm, margins entire, repand, or shallowly dentate.

Cauline leaves

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

(proximal shortly petiolate, distal sessile);

blade linear to elliptic or obovate, (0.5–)1–3.5(–4.5) cm, margins entire or repand.

Racemes

loose, (elongated).

loose.

Flowers

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

sepals oblong, lanceolate, or elliptic, (3–)3.5–6(–7.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (yellow to orange), suborbicular or obovate, (5–)6.5–8(–11) mm, (narrowing gradually to broad claw, usually widened at base).

Fruiting pedicels

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

(recurved, sigmoid), 5–15 mm.

Fruits

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

(sessile or shortly stipitate), orbicular or obovoid, often slightly compressed, (3.5–)4–6 mm;

valves sparsely pubescent, trichomes sessile and stellate, densely pubescent inside, trichomes simple or branched;

ovules 4–12 per ovary;

style 2–4.5 mm.

Seeds

plump, (suborbicular).

flattened.

2n

= 10, 20.

Physaria grahamii

Physaria tenella

Phenology Flowering May–Jun. Flowering Feb–May.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay Sandy soils, gravel, clayey loam, loose rocky slopes, washes, desert slopes and plains, lava hills, frequently in or near bushes
Elevation 2100-2900 m (6900-9500 ft) (0-)600-1900 m ((0-)2000-6200 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
AZ; CA; NV; UT; Mexico (Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 642. FNA vol. 7, p. 663.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda Lesquerella tenella, Lesquerella gordonii var. sessilis
Name authority C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002)
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