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Graham's twinpod

Rogers Pass or klaus' or divide bladderpod

Habit Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. Perennials; caudex simple; densely pubescent, trichomes (loosely spreading), 3–5-rayed, rays distinct, furcate (with exceptionally long branches).
Stems

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

simple from base, erect to decumbent, (slender), 0.6–1.5 dm.

Basal leaves

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

blades obovate to deltate, 1.5–3(–4) cm, margins entire or outer one with 1 or 2 broad teeth.

Cauline leaves

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

blade oblanceolate to spatulate, 0.6–1.5 cm, margins entire.

Racemes

loose, (elongated).

loose.

Flowers

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

sepals (green-yellow, often tinged with purple), elliptic, 3–4.6 mm;

petals oblanceolate, 6–8 mm (claw expanded).

Fruiting pedicels

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

(sigmoid), 5–9 mm.

Fruits

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

(depressed), broadly obovate, compressed (angustiseptate), 2–4 mm, (apex slightly bilobed to nearly truncate);

valves densely pubescent, trichomes strongly ascending, spreading, long, (appearing fuzzy), pubescent inside;

ovules 4 per ovary;

style 3–4 mm, (pubescent or glabrous).

Seeds

plump, (suborbicular).

flattened.

Physaria grahamii

Physaria klausii

Phenology Flowering May–Jun. Flowering Jul.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay Open gravel slides, solifluction cross-stripes of shale rubble, barren shale-derived soil
Elevation 2100-2900 m (6900-9500 ft) 1200-1900 m (3900-6200 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 642. FNA vol. 7, p. 647.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda Lesquerella klausii
Name authority C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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