Physaria grahamii |
Physaria fendleri |
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Graham's twinpod |
Fendler bladderpod, Fendler's bladderpod |
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Habit | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. | Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). |
Stems | several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm. |
Basal leaves | (outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate. |
Cauline leaves | similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
(shortly petiolate); blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute). |
Racemes | loose, (elongated). |
loose to somewhat dense. |
Flowers | sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate); petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short). |
Fruiting pedicels | (ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm. |
Fruits | didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute); valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout; replum as wide as or wider than fruit; ovules (12–)20–32(–40) per ovary; style (2–)3–6 mm. |
Seeds | plump, (suborbicular). |
flattened. |
2n | = 12, 14, 24. |
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Physaria grahamii |
Physaria fendleri |
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Phenology | Flowering May–Jun. | Flowering Mar–May. |
Habitat | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay | Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas |
Elevation | 2100-2900 m (6900-9500 ft) | 100-2000 m (300-6600 ft) |
Distribution |
UT |
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
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Discussion | Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In dry areas, Physaria fendleri may flower following suitable rains at any time of the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 642. | FNA vol. 7, p. 636. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda | Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera |
Name authority | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
Web links |