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Graham's twinpod

Fendler bladderpod, Fendler's bladderpod

Habit Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center).
Stems

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm.

Basal leaves

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate.

Cauline leaves

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

(shortly petiolate);

blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute).

Racemes

loose, (elongated).

loose to somewhat dense.

Flowers

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate);

petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short).

Fruiting pedicels

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm.

Fruits

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute);

valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout;

replum as wide as or wider than fruit;

ovules (12–)20–32(–40) per ovary;

style (2–)3–6 mm.

Seeds

plump, (suborbicular).

flattened.

2n

= 12, 14, 24.

Physaria grahamii

Physaria fendleri

Phenology Flowering May–Jun. Flowering Mar–May.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas
Elevation 2100-2900 m (6900-9500 ft) 100-2000 m (300-6600 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
[WildflowerSearch map]
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In dry areas, Physaria fendleri may flower following suitable rains at any time of the year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 642. FNA vol. 7, p. 636.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera
Name authority C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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