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Graham's twinpod

basin bladderpod

Habit Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. Perennials; caudex branched, (woody); densely pubescent, trichomes (appressed, except spreading on pedicels and fruits, sessile or short-stalked), several-rayed, rays furcate or bifurcate, (strongly tuberculate throughout).
Stems

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

few from base, erect, (stout), to 1.5 dm (sometimes greatly reduced).

Basal leaves

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

blade suborbicular to rhombic or broadly elliptic, 1–4.5 cm, margins entire, (apex rounded to subacute).

Cauline leaves

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

(proximal petiolate, distal subsessile);

blade elliptic, 1–4 cm, (distal narrower), margins entire or remotely dentate.

Racemes

loose, (elongated).

condensed, (subcorymbose to subumbellate, few-flowered).

Flowers

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

sepals (persistent), broadly ovate or oblong to narrowly elliptic, 5.5–8(–9.5) mm, (lateral pair slightly cucullate, median pair thickened apically, cucullate, usually keeled);

petals (orange to yellow), oblong to obovate, 8–11.5(–14.5) mm, (slightly narrowed to broad claw, margins sinuate, often retuse).

Fruiting pedicels

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

(horizontal or divaricate-ascending, straight or slightly curved), 5–15 mm, (stout and rigid).

Fruits

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

(sessile or substipitate), globose, ovoid, or suborbicular, compressed (with marginal and apical constriction), 4–7 mm;

valves pubescent, trichomes contiguous or overlapping, often spreading;

ovules 16–24 per ovary;

style 2–4 mm.

Seeds

plump, (suborbicular).

lenticular, ovate in outline.

2n

= 10.

Physaria grahamii

Physaria cinerea

Phenology Flowering May–Jun. Flowering Mar–May.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay Red soil, chiprock, gypsum or chalky knolls, limestone rubble
Elevation 2100-2900 m (6900-9500 ft) 900-2200 m (3000-7200 ft)
Distribution
from FNA
UT
[BONAP county map]
from FNA
AZ
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 642. FNA vol. 7, p. 631.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda Lesquerella cinerea
Name authority C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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