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spreading bladderpod

Pryor Mountains bladderpod

Habit Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). Perennials; (delicate, short-lived); caudex simple, (sometimes elongated, covered with persistent leaf bases); usually sparsely pubescent, trichomes 7–12-rayed, rays furcate near base.
Stems

simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm.

simple from base, erect to decumbent, (unbranched, mostly filiform, slender), 1–1.5 dm.

Basal leaves

blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent).

(erect, petiole slender);

blades broadly ovate to elliptic, 0.5–1 cm, (base abruptly narrowing to petiole), margins entire.

Cauline leaves

(proximal petiolate, distal sessile);

blade oblanceolate to oblong, 1–7 cm, margins dentate to repand.

(remote, distally shortly petiolate);

blade ± spatulate, (base cuneate), margins entire.

Racemes

loose, (elongated).

lax, (elongated, few-flowered).

Flowers

sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate);

petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw).

sepals (erect), oblong, 3.5–4 mm, (lateral pair not saccate);

petals (often fading to light purple apically), spatulate to nearly lingulate, 6–7 mm.

Fruiting pedicels

(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout).

(recurved to widely spreading, filiform, slender), 5–10 mm.

Fruits

(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm;

valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules 8–20(–28) per ovary;

style 2–4.5 mm.

(pendent), globose or subglobose, compressed, 3–4 mm;

valves ± densely pubescent;

ovules 6–10 per ovary;

style ca. 1.5 mm.

Seeds

slightly flattened.

not seen.

Physaria gracilis

Physaria lesicii

Phenology Flowering Jun(-early Jul).
Habitat Pryor Mountains, on limestone soils in woodlands of Rocky Mountain juniper and/or mountain mahogany, and widely scattered Douglas-fir, fellfields dominated by bluebunch wheatgrass and cushion plants
Elevation 1600-2000 m (5200-6600 ft)
Distribution
from FNA
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
MT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded.
subsp. gracilis
1. Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate.
subsp. nuttallii
Source FNA vol. 7, p. 641. FNA vol. 7, p. 649.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. gracilis subsp. gracilis, P. gracilis subsp. nuttallii
Synonyms Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis Lesquerella lesicii
Name authority (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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