Flora of North America species comparison

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Physaria douglasii subsp. tuplashensis

Physaria douglasii
Photo is of parent taxon

white bluffs bladderpod

Columbia bladder-pod, Douglas' bladder-pod
Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout).
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.
Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.
Cauline leaves

imbricate, blade sometimes orbicular.

similar to basal, blade narrowly linear or, sometimes, orbicular.
Racemes

loose (lax).
Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.
Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.
Fruit(s)

valves: trichomes stalked.

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.
Seeds

flattened.
2n

= 10, 30.

Physaria douglasii subsp. tuplashensis

Physaria douglasii
Phenology Flowering May–Jun(-Aug).
Habitat Caliche soil with sagebrush
Elevation 300 m (1000 ft)
Distribution
from FNA
WA
[BONAP county map]
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

It is possible that subsp. tuplashensis is simply an ecotype, or that its phenotype is in response to its severe habitat on the White Bluffs of the Columbia River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 635. FNA vol. 7, p. 634.
Parent taxa Brassicaceae > tribe Physarieae > Physaria > Physaria douglasii Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. douglasii subsp. douglasii
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella tuplashensis Lesquerella douglasii
Name authority (Rollins: Novon 12: 322. (2002) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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