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Columbia bladder-pod, Douglas' bladder-pod

Snake Range bladderpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Perennials; caudex simple; densely pubescent, trichomes 5–7-rayed, rays bifurcate or trifurcate.
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

several from base, erect or outer usually decumbent toward base, (from below a terminal tuft of mostly erect leaves, unbranched), 1–2 dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

blade ovate to elliptic, 2–4 cm, (base gradually narrowed to petiole), margins entire, (surfaces densely pubescent with trichome layers).

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

(remote, proximal shortly petiolate);

blade spatulate to oblanceolate, similar to basal, (base often cuneate), margins entire.

Racemes

loose (lax).

elongated.

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals linear-oblong, 5–7 mm, (lateral pair subsaccate);

petals (erect), lingulate, 8–10 mm, (claw barely differentiated from blade).

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(recurved), 7–10 mm.

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

(usually pendent, sessile), subglobose, slightly flattened (angustiseptate), 4–5 mm;

valves densely pubescent, trichomes somewhat spreading;

ovules 8–12 per ovary;

style 4–5 mm.

Seeds

flattened.

plump, (mucilaginous when wetted).

2n

= 10, 30.

Physaria douglasii

Physaria pendula

Phenology Flowering May–Jun.
Habitat Limestone gravel and cobbles, typically with junipers
Elevation 1700-2100 m (5600-6900 ft)
Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 634. FNA vol. 7, p. 657.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella douglasii Lesquerella pendula
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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