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Columbia bladder-pod, Douglas' bladder-pod

frosty bladderpod, Piceance bladderpod, Picenace bladderpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Perennials; caudex simple or branched; densely (silvery) pubescent, trichomes (irregularly radiate), 6–8-rayed, rays furcate or bifurcate, fused at base.
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

several from base, prostrate to decumbent, (usually unbranched, rarely branched distally), 1–3 dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

(tufted);

blade broadly obovate, 1–2 cm, margins entire or with 1 or 2 broad teeth, (apex rounded to obtuse).

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

blade oblanceolate to nearly oblong, similar to basal, (base cuneate), margins entire.

Racemes

loose (lax).

(secund), loose, (elongated in fruit).

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals (yellowish), elliptic to lanceolate, (2–)3–4 mm;

petals spatulate, (3.9–)5–7 mm.

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(recurved), 6–8(–12) mm.

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

(usually pendent), elliptic to subglobose, usually slightly compressed (latiseptate), 3–4 mm;

valves densely pubescent, sometimes with scattered trichomes inside;

ovules 4 per ovary;

style ca. 3 mm.

Seeds

flattened.

somewhat flattened.

2n

= 10, 30.

Physaria douglasii

Physaria parviflora

Phenology Flowering Jun–Jul.
Habitat Shale of steep slopes, rock crevices, ledges, canyon sides, shale-marlstone
Elevation 2100-2700 m (6900-8900 ft)
Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria parviflora is known from the Parachute Creek Member of the Green River Formation, Rio Blanco County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 634. FNA vol. 7, p. 657.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella douglasii Lesquerella parviflora
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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