Physaria douglasii |
Physaria pallida |
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Columbia bladder-pod, Douglas' bladder-pod |
white bladderpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Annuals (winter); with a fine taproot; sparsely pubescent, trichomes (minute), 3- or 4-rayed, rays furcate or, sometimes, trifurcate. | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
few to several from base, erect, (from basal leaf cluster, branched distally, flowering branches slender, subtended by bracts), 3–6 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
blade oblanceolate or broadly obovate, to 10 cm, margins usually sinuate-dentate or entire, sometimes lobed. |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
(proximal shortly petiolate, distal sessile); blade oblanceolate to narrowly oblong, similar to basal, (distal with base slightly cuneate). |
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Racemes | loose (lax). |
paniculate, (rachises and pedicels more densely pubescent than proximal leaves). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals elliptic, 3–7 mm, (median pair slightly thickened apically, cucullate); petals (white), broadly ovate, to 12 mm, (narrowing gradually to short claw). |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(widely divaricate-ascending and straight, or slightly recurved), 10–15 mm, (slender, pubescent). |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(widely spreading to nearly pendent in age, shortly stipitate), globose or subglobose, not or slightly inflated, 3–10 mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–12 per ovary; style ca. 2 mm, (slender, fragile). |
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Seeds | flattened. |
flattened. |
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2n | = 10, 30. |
= 12. |
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Physaria douglasii |
Physaria pallida |
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Phenology | Flowering Apr–May. | |||||
Habitat | Grassy openings of small glade prairies, outcrops | |||||
Elevation | 90 m (300 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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TX |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pallida is known from the Weches Formation in San Augustine County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 656. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | Vesicaria grandiflora var. pallida, Alyssum pallidum, Lesquerella pallida, Vesicaria pallida | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |
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