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Columbia bladder-pod, Douglas' bladder-pod

white bladderpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Annuals (winter); with a fine taproot; sparsely pubescent, trichomes (minute), 3- or 4-rayed, rays furcate or, sometimes, trifurcate.
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

few to several from base, erect, (from basal leaf cluster, branched distally, flowering branches slender, subtended by bracts), 3–6 dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

blade oblanceolate or broadly obovate, to 10 cm, margins usually sinuate-dentate or entire, sometimes lobed.

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

(proximal shortly petiolate, distal sessile);

blade oblanceolate to narrowly oblong, similar to basal, (distal with base slightly cuneate).

Racemes

loose (lax).

paniculate, (rachises and pedicels more densely pubescent than proximal leaves).

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals elliptic, 3–7 mm, (median pair slightly thickened apically, cucullate);

petals (white), broadly ovate, to 12 mm, (narrowing gradually to short claw).

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(widely divaricate-ascending and straight, or slightly recurved), 10–15 mm, (slender, pubescent).

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

(widely spreading to nearly pendent in age, shortly stipitate), globose or subglobose, not or slightly inflated, 3–10 mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–12 per ovary;

style ca. 2 mm, (slender, fragile).

Seeds

flattened.

flattened.

2n

= 10, 30.

= 12.

Physaria douglasii

Physaria pallida

Phenology Flowering Apr–May.
Habitat Grassy openings of small glade prairies, outcrops
Elevation 90 m (300 ft)
Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria pallida is known from the Weches Formation in San Augustine County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 634. FNA vol. 7, p. 656.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella douglasii Vesicaria grandiflora var. pallida, Alyssum pallidum, Lesquerella pallida, Vesicaria pallida
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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