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Columbia bladder-pod, Douglas' bladder-pod

double bladderpod, double twinpod

Habit Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout).
Stems

simple from base, erect, (usually unbranched), to 4.5 dm.

several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm.

Basal leaves

blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid.

(petiole somewhat winged);

blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy).

Cauline leaves

similar to basal, blade narrowly linear or, sometimes, orbicular.

blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute).

Racemes

loose (lax).

moderately dense (or elongated).

Flowers

sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate);

petals 6–11 mm.

sepals linear-oblong, 6–8 mm;

petals spatulate, 9–12 mm.

Fruiting pedicels

(recurved, straight, curved, or sigmoid), 6–20 mm.

(divergent, straight to somewhat curved or sigmoid), 5–12 mm.

Fruits

obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm;

valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked;

ovules 4(–8) per ovary;

style (1.6–)3–6 mm.

(erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad);

valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading;

replum linear-oblong, constricted, as wide as or wider than fruit;

ovules 4 per ovary;

style 4–5(–9) mm.

Seeds

flattened.

plump, (broad).

2n

= 10, 30.

= 8, 16.

Physaria douglasii

Physaria brassicoides

Phenology Flowering May–Jun.
Habitat Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks
Elevation 900-1400 m (3000-4600 ft)
Distribution
from FNA
ID; MT; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; ND; NE; SD; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruit valves: trichomes sessile; cauline leaves loosely arranged, blades narrowly linear; British Columbia, Idaho, Montana, Oregon, Washington.
subsp. douglasii
1. Fruit valves: trichomes stalked; cauline leaves imbricate, blades sometimes orbicular; White Bluffs adjacent to Columbia River of Washington.
subsp. tuplashensis
Source FNA vol. 7, p. 634. FNA vol. 7, p. 628.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. douglasii subsp. douglasii, P. douglasii subsp. tuplashensis
Synonyms Lesquerella douglasii
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Rydberg: Bull. Torrey Bot. Club 29: 237. (1902)
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