Physaria douglasii |
Physaria brassicoides |
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Columbia bladder-pod, Douglas' bladder-pod |
double bladderpod, double twinpod |
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Habit | Perennials; caudex simple; densely pubescent, trichomes (sessile or nearly so), 4–6(–10)-rayed, rays usually furcate near base, rarely bifurcate, (umbonate, tuberculate throughout). | Perennials; (somewhat compact); caudex branched, (relatively large); (silvery) pubescent throughout, trichomes (sessile), several-rayed, rays furcate, (slightly umbonate, tuberculate throughout). | ||||
Stems | simple from base, erect, (usually unbranched), to 4.5 dm. |
several from base, decumbent to ascending (arising laterally, unbranched, stout), (0.2–)0.5–1.7 dm. |
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Basal leaves | blade suborbicular to elliptic, 2–9.5(–11.5) cm, margins entire, sinuate, coarsely dentate, or almost lyrate-pinnatifid. |
(petiole somewhat winged); blades orbicular to obovate, 2–6 cm (width 1–2.5 cm, thick), margins usually repand, rarely entire, (adaxial surface scurfy). |
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Cauline leaves | similar to basal, blade narrowly linear or, sometimes, orbicular. |
blade oblanceolate to broadly spatulate, 1–2 cm (width 3–5 mm), margins entire, (apex obtuse to subacute). |
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Racemes | loose (lax). |
moderately dense (or elongated). |
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Flowers | sepals elliptic or ovate, (2–)3.5–7.5 mm, (cucullate); petals 6–11 mm. |
sepals linear-oblong, 6–8 mm; petals spatulate, 9–12 mm. |
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Fruiting pedicels | (recurved, straight, curved, or sigmoid), 6–20 mm. |
(divergent, straight to somewhat curved or sigmoid), 5–12 mm. |
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Fruits | obovoid to subglobose, not inflated (not angustiseptate), 3–6 mm; valves sparsely pubescent, sometimes glabrous inside, trichomes sessile or stalked; ovules 4(–8) per ovary; style (1.6–)3–6 mm. |
(erect), didymous, cordate, moderately inflated, (6–)10–20 × 10–23 mm, (papery, base obtuse or with obscure sinus, apical sinus deep, broad); valves (retaining seeds after dehiscence), densely and loosely pubescent, trichomes spreading; replum linear-oblong, constricted, as wide as or wider than fruit; ovules 4 per ovary; style 4–5(–9) mm. |
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Seeds | flattened. |
plump, (broad). |
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2n | = 10, 30. |
= 8, 16. |
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Physaria douglasii |
Physaria brassicoides |
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Phenology | Flowering May–Jun. | |||||
Habitat | Bare hillsides, dry gravel and clay soil, badlands, clay knolls, banks | |||||
Elevation | 900-1400 m (3000-4600 ft) | |||||
Distribution |
ID; MT; OR; WA; BC
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CO; MT; ND; NE; SD; WY
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 634. | FNA vol. 7, p. 628. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella douglasii | |||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | Rydberg: Bull. Torrey Bot. Club 29: 237. (1902) | ||||
Web links |
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