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Idaho twinpod, salmon twinpod

common twinpod, double twinpod, twin bladderpod

Habit Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate).
Stems

several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm.

Basal leaves

blade margins ± lyrate or, rarely, entire.

(forming a strong rosette; long-petiolate);

blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery).

Cauline leaves

blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute).

Trichomes

appressed throughout.

Racemes

congested, (elongated in fruit, greatly exceeding leaves).

Flowers

sepals lanceolate to oblong, 6–8 mm, (often keeled);

petals spatulate, 10–12 mm.

Fruiting pedicels

(spreading, straight or slightly curved), 8–12 mm.

Fruits

highly inflated, (papery);

ovules 8 per ovary.

(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed);

valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy);

replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse;

ovules (4–)8 per ovary;

style 7–9 mm.

Seeds

flattened.

2n

= 24.

Physaria didymocarpa subsp. lyrata

Physaria didymocarpa

Phenology Flowering May–Jun.
Habitat Gravel, alluvial fans, steep banks
Elevation 1300-2100 m (4300-6900 ft)
Distribution
from FNA
ID
[BONAP county map]
from FNA
ID; MT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Subspecies lyrata is found only in the Salmon River drainage.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trichomes spreading throughout; basal leaf blade margins dentate.
subsp. lanata
1. Trichomes appressed (except fruits in subsp. didymocarpa); basal leaf blade margins repand to dentate, ± lyrate or, rarely, entire
→ 2
2. Fruits moderately inflated, trichomes spreading.
subsp. didymocarpa
2. Fruits highly inflated (papery), trichomes appressed.
subsp. lyrata
Source FNA vol. 7, p. 634. FNA vol. 7, p. 633.
Parent taxa Brassicaceae > tribe Physarieae > Physaria > Physaria didymocarpa Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. didymocarpa subsp. didymocarpa, P. didymocarpa subsp. lanata
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. didymocarpa subsp. didymocarpa, P. didymocarpa subsp. lanata, P. didymocarpa subsp. lyrata
Synonyms P. didymocarpa var. lyrata Vesicaria didymocarpa, Coulterina didymocarpa
Name authority (C. L. Hitchcock) O’Kane: Novon 17: 379. (2007) (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848)
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