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Dudley bluffs bladderpod

silver twinpod, west silver bladderpod

Habit Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). Perennials; (diminutive); caudex simple or branched, (buried, with thatch of persistent leaf bases distally); (appearing silvery gray-green to silvery purple), densely pubescent, trichomes usually 5 or 6 (rarely 7)-rayed, rays bifurcate or incompletely so, (relatively short, stout, umbonate, moderately tuberculate to nearly smooth, lower layer smoother).
Stems

simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm.

1–5 from base, prostrate to slightly decumbent, (arising laterally, also erect or ascending from tuft of basal leaves, unbranched, purple-green), 0.08–0.3 dm.

Basal leaves

similar to cauline, (erect, surfaces silvery).

(petiole slightly winged);

blade oblanceolate, elliptic, or rhombic, (mostly flat, sometimes somewhat folded), 0.6–2.7 cm, (base tapering to petiole), margins entire, (apex rounded to rounded-acute).

Cauline leaves

(ascending, subsessile);

blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse).

(3–7, shortly petiolate or sessile);

blade elliptic to oblanceolate, 0.3–0.5 cm, margins entire.

Racemes

strongly congested, (often sessile or nearly so, lateral to leaves).

crowded, (ca. 3–7 fruits).

Flowers

sepals (loosely erect), narrowly oblong, 3–4 mm;

petals spatulate, 5–6 mm.

sepals (greenish yellow), linear-triangular, 3.7–5 mm, (lateral pair subsaccate);

petals oblanceolate to narrowly obovate, 4.5–9 mm.

Fruiting pedicels

(erect or ascending, straight to slightly curved), 3–6 mm.

(ascending, straight), 1.8–3.4 mm.

Fruits

ovate, compressed (latiseptate) on margins and apically, 4–5 mm;

valves pubescent, trichomes densely appressed;

ovules 4 per ovary;

style 1–1.5 mm.

(shortly stipitate, purple or greenish purple in age), slightly didymous, ovoid to obpyriform, 3–5 mm (wider than long, base rounded-obtuse, apex rounded, flattened, or slightly emarginate);

valves (inflated, slightly wider than replum), pubescent, trichomes scattered;

replum obovate to orbicular-obdeltate, apex rounded, obtuse, or truncate;

septum complete or medially small-perforate;

ovules 4–6(–8) per ovary;

style 2–3.6 mm.

Seeds

plump.

relatively plump, (ovate to suborbicular, usually rounded on one side, ± flat or concave on the other, not mucilaginous when wetted).

Physaria congesta

Physaria scrotiformis

Phenology Flowering Apr–May. Flowering Jun-early Jul.
Habitat Barren knolls with pinyon-juniper Tundra areas with islands of Engelmann spruce on Leadville limestone, amidst limestone cobbles and gravel
Elevation 1800-2100 m (5900-6900 ft) 3500-3700 m (11500-12100 ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
CO
[BONAP county map]
Discussion

Of conservation concern.

Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria scrotiformis is known only from La Plata County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 632. FNA vol. 7, p. 662.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella congesta
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) O’Kane: Novon 17: 376, fig. 1. (2007)
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