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Dudley bluffs bladderpod

Fremont County twinpod, Rocky Mountain or Fremont County twinpod

Habit Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). Perennials; caudex usually simple; (silvery) pubescent throughout, trichome rays furcate.
Stems

simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm.

several from base, prostrate to decumbent, 0.3–1 dm.

Basal leaves

similar to cauline, (erect, surfaces silvery).

(rosulate; petiole winged);

blade orbicular to broadly obovate, 1.5–3 cm, margins entire or with broad, obscure toothlike angles each side at apex, (apex obtuse, surfaces densely pubescent, trichomes appressed).

Cauline leaves

(ascending, subsessile);

blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse).

blade broadly spatulate to linear-oblanceolate, 1–1.5 cm, margins entire.

Racemes

strongly congested, (often sessile or nearly so, lateral to leaves).

condensed, (subumbellate to slightly more elongated, few-flowered).

Flowers

sepals (loosely erect), narrowly oblong, 3–4 mm;

petals spatulate, 5–6 mm.

sepals (yellowish, often with some purple), narrowly lanceolate, 5–6 mm;

petals spatulate, 7.3–9.2 mm, (not clawed).

Fruiting pedicels

(erect or ascending, straight to slightly curved), 3–6 mm.

(divaricate-ascending, straight to slightly curved), 6–10 mm.

Fruits

ovate, compressed (latiseptate) on margins and apically, 4–5 mm;

valves pubescent, trichomes densely appressed;

ovules 4 per ovary;

style 1–1.5 mm.

didymous, irregular, suborbicular, deeply bilobed, inflated in age, 10–12 × 12–15 mm, (papery, basal sinus absent or obsolete, apical sinus deep);

valves (retaining seeds after dehiscence), densely pubescent, trichomes spreading, (ovaries and immature fruit downy);

replum narrowly ovate to broadly oblong, not narrowed at middle, as wide as or wider than fruit, apex acute to obtuse;

ovules 4 per ovary;

style 3–7 mm.

Seeds

plump.

flattened.

Physaria congesta

Physaria saximontana

Phenology Flowering Apr–May.
Habitat Barren knolls with pinyon-juniper
Elevation 1800-2100 m (5900-6900 ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
MT; WY
[BONAP county map]
Discussion

Of conservation concern.

Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Physaria saximontana (especially subsp. dentata) is morphologically similar to 22. P. didymocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaf blades: margins entire, apices rounded to angled; styles 3-5 mm.
subsp. saximontana
1. Basal leaf blades: margins dentate (teeth broad), apices angled; styles 4-7 mm.
subsp. dentata
Source FNA vol. 7, p. 632. FNA vol. 7, p. 661.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. saximontana subsp. dentata, P. saximontana subsp. saximontana
Synonyms Lesquerella congesta
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) Rollins: Contr. Gray Herb. 214: 13. (1984)
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