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Dudley bluffs bladderpod

alpine bladderpod, reed's twinpod, Rollins' bladderpod

Habit Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout).
Stems

simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm.

few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm.

Basal leaves

similar to cauline, (erect, surfaces silvery).

(erect);

blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire.

Cauline leaves

(ascending, subsessile);

blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse).

blade linear, similar to basal.

Racemes

strongly congested, (often sessile or nearly so, lateral to leaves).

dense, (often subumbellate, not or barely exceeding basal leaves).

Flowers

sepals (loosely erect), narrowly oblong, 3–4 mm;

petals spatulate, 5–6 mm.

sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals lingulate, 6–9 mm.

Fruiting pedicels

(erect or ascending, straight to slightly curved), 3–6 mm.

(ascending, curved), 3–5.5 mm.

Fruits

ovate, compressed (latiseptate) on margins and apically, 4–5 mm;

valves pubescent, trichomes densely appressed;

ovules 4 per ovary;

style 1–1.5 mm.

lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm;

valves pubescent, trichomes closely appressed;

ovules 8–12 per ovary;

style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma).

Seeds

plump.

plump, (oblong).

Physaria congesta

Physaria reediana

Phenology Flowering Apr–May. Flowering May–Jul.
Habitat Barren knolls with pinyon-juniper Open areas of grasslands on calcareous soils
Elevation 1800-2100 m (5900-6900 ft) 1200-1900 m (3900-6200 ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
CO; NE; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 632. FNA vol. 7, p. 660.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella congesta Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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