Physaria congesta |
Physaria ludoviciana |
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Dudley bluffs bladderpod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
Basal leaves | similar to cauline, (erect, surfaces silvery). |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
Cauline leaves | (ascending, subsessile); blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse). |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
Racemes | strongly congested, (often sessile or nearly so, lateral to leaves). |
compact, (elongated and loose in fruit, densely-flowered). |
Flowers | sepals (loosely erect), narrowly oblong, 3–4 mm; petals spatulate, 5–6 mm. |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
Fruiting pedicels | (erect or ascending, straight to slightly curved), 3–6 mm. |
(usually recurved), (5–)10–20(–25) mm. |
Fruits | ovate, compressed (latiseptate) on margins and apically, 4–5 mm; valves pubescent, trichomes densely appressed; ovules 4 per ovary; style 1–1.5 mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
Seeds | plump. |
slightly flattened. |
2n | = 10, 20, 30. |
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Physaria congesta |
Physaria ludoviciana |
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Phenology | Flowering Apr–May. | Flowering Apr–Jul(-Aug). |
Habitat | Barren knolls with pinyon-juniper | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands |
Elevation | 1800-2100 m (5900-6900 ft) | 0-1900 m (0-6200 ft) |
Distribution |
CO |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Of conservation concern. Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 632. | FNA vol. 7, p. 649. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella congesta | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana |
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |