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Dudley bluffs bladderpod

foothill bladderpod, silver bladderpod

Habit Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout).
Stems

simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm.

few from base, erect with outer usually decumbent, 1–3.5(–5) dm.

Basal leaves

similar to cauline, (erect, surfaces silvery).

(erect);

blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes).

Cauline leaves

(ascending, subsessile);

blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse).

blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute.

Racemes

strongly congested, (often sessile or nearly so, lateral to leaves).

compact, (elongated and loose in fruit, densely-flowered).

Flowers

sepals (loosely erect), narrowly oblong, 3–4 mm;

petals spatulate, 5–6 mm.

sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate);

petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw).

Fruiting pedicels

(erect or ascending, straight to slightly curved), 3–6 mm.

(usually recurved), (5–)10–20(–25) mm.

Fruits

ovate, compressed (latiseptate) on margins and apically, 4–5 mm;

valves pubescent, trichomes densely appressed;

ovules 4 per ovary;

style 1–1.5 mm.

subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm;

valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile;

ovules (4–)8–12(–16) per ovary;

style 3–4.5(–6.5) mm.

Seeds

plump.

slightly flattened.

2n

= 10, 20, 30.

Physaria congesta

Physaria ludoviciana

Phenology Flowering Apr–May. Flowering Apr–Jul(-Aug).
Habitat Barren knolls with pinyon-juniper Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands
Elevation 1800-2100 m (5900-6900 ft) 0-1900 m (0-6200 ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 632. FNA vol. 7, p. 649.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella congesta Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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