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Dudley bluffs bladderpod

Idaho bladderpod

Habit Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). Perennials; caudex simple, (often enlarged by persistent leaf bases); densely pubescent, trichomes (sessile or short-stalked), rays furcate or bifurcate, (nearly smooth to finely tuberculate).
Stems

simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm.

simple from base, decumbent, (occasionally few-branched), 0.5–1.5(–2) dm.

Basal leaves

similar to cauline, (erect, surfaces silvery).

blade elliptic to broadly obovate, triangular, rhombic, or round, 1.5–3(–4) cm, margins often sinuate or shallowly lobed.

Cauline leaves

(ascending, subsessile);

blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse).

(sessile or shortly petiolate);

blade elliptic to oblanceolate to obovate, 0.5–1.5 cm, (base narrowed to petiole), margins entire.

Racemes

strongly congested, (often sessile or nearly so, lateral to leaves).

compact to loose.

Flowers

sepals (loosely erect), narrowly oblong, 3–4 mm;

petals spatulate, 5–6 mm.

sepals oblong to broadly elliptic, 4–7.5 mm, (lateral pair saccate or not);

petals spatulate, 7.5–10 mm.

Fruiting pedicels

(erect or ascending, straight to slightly curved), 3–6 mm.

(ascending or divaricate-spreading, straight to loosely sigmoid or curved), 4–10 mm.

Fruits

ovate, compressed (latiseptate) on margins and apically, 4–5 mm;

valves pubescent, trichomes densely appressed;

ovules 4 per ovary;

style 1–1.5 mm.

(sessile or substipitate), elliptic, suborbicular, or oblong, strongly compressed (angustiseptate), 5–9 mm, (rounded to sharply keeled on 1 side, edges ± keeled);

valves: (margins covering replum edges or not), usually pubescent throughout or, rarely, glabrous inside;

ovules (4–)8–14(–16) per ovary;

style 2–4.5(–5) mm.

Seeds

plump.

slightly flattened.

Physaria congesta

Physaria carinata

Phenology Flowering Apr–May.
Habitat Barren knolls with pinyon-juniper
Elevation 1800-2100 m (5900-6900 ft)
Distribution
from FNA
CO
[BONAP county map]
from FNA
ID; MT; WY
[BONAP county map]
Discussion

Of conservation concern.

Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Differences in fruit morphology become blurred and the three subspecies are often indistinguishable where their ranges meet near the intersection of Idaho, Montana, and Wyoming.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits elliptic, not keeled, valve margins (thin, rounded), not covering replum edges.
subsp. paysonii
1. Fruits elliptic, suborbicular, or broadly oblong, keeled on one side, valve margins covering replum edges
→ 2
2. Fruits elliptic, bases narrow-rounded to acute, apices ± acute.
subsp. carinata
2. Fruits suborbicular to elliptic or broadly oblong, bases and apices rounded.
subsp. pulchella
Source FNA vol. 7, p. 632. FNA vol. 7, p. 629.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. carinata subsp. carinata, P. carinata subsp. paysonii, P. carinata subsp. pulchella
Synonyms Lesquerella congesta Lesquerella carinata
Name authority (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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