Physaria congesta |
Physaria acutifolia |
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Dudley bluffs bladderpod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; (relatively diminutive, strongly condensed); caudex (buried), simple or branched, (stout, thatched, thickened with persistent leaf bases); densely pubescent, trichomes (appressed, stiff), 4- or 5-rayed, rays fused at center, (mostly bifurcate). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). |
Stems | simple or few from base, decumbent to ascending, (arising laterally from a tight hemispherical tuft of leaves), to 0.15 dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
Basal leaves | similar to cauline, (erect, surfaces silvery). |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
Cauline leaves | (ascending, subsessile); blade linear-oblanceolate, (0.6–)0.8–1.3(–1.5) cm, margins entire, (apex acute to narrowly obtuse). |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
Racemes | strongly congested, (often sessile or nearly so, lateral to leaves). |
loose, (elongated in fruit). |
Flowers | sepals (loosely erect), narrowly oblong, 3–4 mm; petals spatulate, 5–6 mm. |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
Fruiting pedicels | (erect or ascending, straight to slightly curved), 3–6 mm. |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
Fruits | ovate, compressed (latiseptate) on margins and apically, 4–5 mm; valves pubescent, trichomes densely appressed; ovules 4 per ovary; style 1–1.5 mm. |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
Seeds | plump. |
(dark brown), flattened, (2–3 mm). |
2n | = 10, 16, 24. |
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Physaria congesta |
Physaria acutifolia |
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Phenology | Flowering Apr–May. | Flowering May–Jun(-Jul). |
Habitat | Barren knolls with pinyon-juniper | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities |
Elevation | 1800-2100 m (5900-6900 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
CO |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Of conservation concern. Physaria congesta is found on white, decomposed shale of the Thirteenmile Creek Tongue of the Green River Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 632. | FNA vol. 7, p. 624. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella congesta | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa |
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) |
Web links |