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Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

Snake Range bladderpod

Habit Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small). Perennials; caudex simple; densely pubescent, trichomes 5–7-rayed, rays bifurcate or trifurcate.
Stems

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

several from base, erect or outer usually decumbent toward base, (from below a terminal tuft of mostly erect leaves, unbranched), 1–2 dm.

Basal leaves

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

blade ovate to elliptic, 2–4 cm, (base gradually narrowed to petiole), margins entire, (surfaces densely pubescent with trichome layers).

Cauline leaves

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

(remote, proximal shortly petiolate);

blade spatulate to oblanceolate, similar to basal, (base often cuneate), margins entire.

Racemes

subcorymbose.

elongated.

Flowers

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

sepals linear-oblong, 5–7 mm, (lateral pair subsaccate);

petals (erect), lingulate, 8–10 mm, (claw barely differentiated from blade).

Fruiting pedicels

(divaricate, straight), 5–10 mm.

(recurved), 7–10 mm.

Fruits

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

(usually pendent, sessile), subglobose, slightly flattened (angustiseptate), 4–5 mm;

valves densely pubescent, trichomes somewhat spreading;

ovules 8–12 per ovary;

style 4–5 mm.

Seeds

flattened, (2–3 mm).

plump, (mucilaginous when wetted).

2n

= 48–52, 52, 64, 67–70.

Physaria alpestris

Physaria pendula

Phenology Flowering May–Jun. Flowering May–Jun.
Habitat Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities Limestone gravel and cobbles, typically with junipers
Elevation (700-)1300-2400 m ((2300-)4300-7900 ft) 1700-2100 m (5600-6900 ft)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 657.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpestris Lesquerella pendula
Name authority Suksdorf: W. Amer. Sci. 15: 58. (1906) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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